Eutitanosauria

Description

Eutitanosauria is a clade of titanosaurs, encompassing the more derived members of the group and characterized by the absence of the hyposphene-hypantrum articulation and possibly the presence of osteoderms. The group was first named by Sanz and colleagues in 1999, who used it to unite the group of Argyrosaurus, Lirainosaurus, Saltasaurus and the Peiropolis titanosaur. However, this definition was not used as it made the group equivalent to Saltasauridae, so Saldago redefined it in 2003 to be all titanosaurs closer to Saltasaurus than Epachthosaurus. This definition created Eutitanosauria as the sister group to Epachthosaurinae (Epachthosaurus but not Saltasaurus), but was problematic due to the variable nature of Epachthosaurus. Eutitanosauria was often broadly similar to Lithostrotia, and has often been unused or unlabelled on phylogenies. Sometimes Epachthosaurus would be more primitive than Malawisaurus, making Eutitanosauria more encompassing than Lithostrotia, or Epachthosaurus could nest close to Colossosauria and limit Eutitanosauria to a smaller group of saltasauroids. Because of the flexible nature of Epachthosaurus in basal titanosaur phylogeny, Carballido and colleagues redefined the group in 2022 to include the smallest clade of both Patagotitan, a colossosaur, and Saltasaurus, creating a node-stem clade with Colossosauria and Saltasauroidea, presenting the informal cladogram of stable titanosaur clades below.

Information(s)

Source: The Paleobiology Database

Attibution: ?
Statut: Valide
Environnement de découverte: terrestrial
Mode de vie: terrestrial
Mode de locomotion: actively mobile
Vision: ?
Alimentation: herbivore
Mode de reprodution: oviparous, dispersal=direct/internal,mobile
Classification: Titanosauria >> Titanosauriformes >> Macronaria >> Neosauropoda >> Eusauropoda >> Gravisauria >> Sauropoda >> Saurischia >> Dinosauria
Période: Albian - Maastrichtian (de -113.00 Ma à -66.00 Ma)

Descendance(s):

Genres: Argentinosaurus Bonitasaura Dreadnoughtus Inawentu Lohuecotitan Magyarosaurus Menucocelsior Normanniasaurus Paludititan Patagotitan Petrustitan

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Lirainosaurinae: Ampelosaurus Atsinganosaurus Garrigatitan Lirainosaurus

Découverte(s): 44 occcurrences
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- Argentine
  - Chubut
    - ?
      - Formation Cerro Barcino
        Patagotitan mayorum⁷⁶³¹⁹
  - Neuquén
    - ?
      - Formation Bajo de la Carpa
        Inawentu oslatus⁸⁶³⁵⁵
      - Formation Huincul
        Argentinosaurus huinculensis¹³⁹⁰¹
  - Río Negro
    - Cerro Policía
      - Formation Bajo de la Carpa
        Bonitasaura salgadoi¹¹⁸²⁰
    - El Cuy
      - Formation Allen
        Menucocelsior arriagadai⁷⁸²⁶⁵
      - Formation Bajo de la Carpa
        Bonitasaura⁷⁶³⁰⁴
  - Santa Cruz
    - ?
      - Formation Cerro Fortaleza
        Dreadnoughtus schrani⁸⁶⁷²⁵
        Dreadnoughtus schrani⁵²⁵⁷⁷
- Espagne
  - Castilla y Leon
    - Burgos
      - Formation Vitoria
        Lirainosaurus astibiae⁹⁵⁵⁹
        Lirainosaurus astibiae⁹⁵⁵⁹
        Lirainosaurus astibiae⁹⁵⁵⁹
  - Castilla-La Mancha
    - Cuenca
      - Formation Villalba de la Sierra
        Ampelosaurus⁴⁴⁴²¹
        Lohuecotitan pandafilandi⁶⁰⁰⁸⁸
    - Guadalajara
      - Formation Villalba de la Sierra
        Lirainosaurus⁷⁶¹²⁹
  - Valencia
    - ?
      - Formation Sierra Perenchiza
        Lirainosaurus astibiae³²⁷⁴⁹
        Lirainosaurus astibiae³²⁷⁴⁹
        Lirainosaurus astibiae³²⁷⁴⁹
        Lirainosaurus astibiae³²⁷⁴⁹
- France
  - Languedoc-Roussillon
    - Aude
      - Formation Marnes Rouges Inférieures
        Ampelosaurus atacis¹³⁰¹⁹
        Lirainosaurus astibiae⁵⁵⁴²⁹
    - Hérault
      - Formation Grès à Reptiles
        Ampelosaurus atacis¹³⁰³²
  - Provence-Alpes-Cote d'Azur
    - Bouches-du-Rhône
      - Formation ?
        Ampelosaurus atacis⁵⁵⁴²⁹
  - Provence-Alpes-Côte d'Azur
    - ?
      - Formation ?
        Garrigatitan meridionalis⁷⁴⁸⁹⁰
    - Bouches-du-Rhône
      - Formation ?
        Atsinganosaurus velauciensis³³¹⁹⁷
    - Var
      - Formation Argiles rutilantes
        Ampelosaurus atacis⁵⁵⁴²⁹
      - Formation Grès à Reptiles
        Lirainosaurus astibiae⁵⁵⁴²⁹
  - Seine-Maritime
    - ?
      - Formation Poudingue ferrugineux
        Normanniasaurus genceyi⁴⁸⁴⁷¹
        Normanniasaurus genceyi⁴⁸⁴⁷¹
- Roumanie
  - Alba
    - ?
      - Formation Sebeş
        Magyarosaurus⁴⁹⁵⁵⁷
        Magyarosaurus⁴⁹⁵⁵⁷
        Magyarosaurus⁴⁹⁵⁵⁷
        Magyarosaurus⁴⁹⁵⁵⁷
        Petrustitan hungaricus identifié comme Magyarosaurus ? hungaricus⁸¹⁰³¹
      - Formation Şard
        Magyarosaurus dacus²⁷¹⁸³
  - Hunedoara
    - ?
      - Formation ?
        Magyarosaurus dacus⁹¹²⁸²
      - Formation Densuş-Ciula
        Magyarosaurus dacus⁹¹²⁸²
        Magyarosaurus dacus¹⁹³⁷⁷
        Magyarosaurus dacus identifié comme Titanosaurus n. sp. dacus³²⁹⁴³
      - Formation Râul Mare
        Paludititan⁷⁶⁰⁹²
        Paludititan nalatzensis³⁷²⁵⁵
      - Formation Sînpetru
        Magyarosaurus dacus¹⁹³⁷⁷
        Magyarosaurus dacus identifié comme Magyarosaurus n. sp. transsylvanicus²⁹⁴⁸³
        Petrustitan hungaricus⁹¹²⁸²
        Petrustitan hungaricus identifié comme Magyarosaurus n. sp. hungaricus²⁹⁴⁸³
Historique des modifications:
- 2025-03-09: Champ(s) mis à jour : Nombre d'occurences Age d'apparition (min) Age de disparition (max)
- 2025-02-01: Champ(s) mis à jour : Rang Nom accepté
- 2024-11-15: Champ(s) mis à jour : Nombre d'occurences
- 2024-09-11: Champ(s) mis à jour : Nombre d'occurences Age de disparition (max)
- 2024-09-07: Création d'une famille à partir des données de pbdb

Publication(s)

La base comprend 28 publication(s).

Source: The Paleobiology Database

↑¹J. L. Carballido, D. Pol, and A. Otero, I. A. Cerda, L. Salgado, A. C. Garrido, J. Ramezzani, N. R. Cúneo, M. J. Krause. 2016. A new giant titanosaur from the Early Cretaceous of Chubut sheds light to giant titanosaur evolution. XXX Jornadas Argentinas de Paleontología de Vertebrados. Resúmenes. Ameghiniana 53(6 (suppl.)):11
↑¹L. S. Filippi, R. D. Juárez Valieri, and P. A. Gallina, A. H. Méndez, F. A. Gianechini, A. C. Garrido. 2024. A rebbachisaurid-mimicking titanosaur and evidence of a Late Cretaceous faunal disturbance event in South-West Gondwana. Cretaceous Research 154:105754 (https://doi.org/10.1016/j.cretres.2023.105754)
↑¹J. F. Bonaparte and R. A. Coria. 1993. Un nuevo y gigantesco saurópodo titanosaurio de la Formación Rio Limay (Albanio-Cenomaniano) de la Provincia del Neuquén, Argentina [A new and huge titanosaur sauropod from the Rio Limay Formation (Albian-Cenomanian) of Neuquén Province, Argentina]. Ameghiniana 30(3):271-282
↑¹S. Apesteguía. 2004. Bonitasaura salgadoi gen. et sp. nov.: a beaked sauropod from the Late Cretaceous of Patagonia. Naturwissenschaften 91(10):493-497
↑¹M. A. Rolando, J. A. Garcia Marsà, and F. L. Agnolín, M. J. Motta, S. Rozadilla, F. E. Novas. 2022. The sauropod record of Salitral Ojo del Agua: A Upper Cretaceous (Allen Formation) fossiliferous locality from northern Patagonia, Argentina. Cretaceous Research 129:105029:1-25 (https://doi.org/10.1016/j.cretres.2021.105029)
↑¹P. A. Gallina and A. Otero. 2015. Reassessment of Laplatasaurus araukanicus (Sauropoda: Titanosauria) from the Upper Cretaceous of Patagonia, Argentina. Ameghiniana 52(5):487-501 (https://doi.org/10.5710/AMGH.08.06.2015.2911)
↑¹F. L. Agnolín, G. Álvarez Herrera, and M. Aranciaga Rolando, M. Motta, S. Rozadilla, L. Verdiquio, J. S. D.'Angelo, D. Moyano-Paz, A. N. Varela, J. Sterli, S. Bogan, S. Miner, A. M. Rodríguez, G. Munoz, M. P. Isasi , F. E. Novas. 2023. Fossil vertebrates from the Cerro Fortaleza Formation (Upper Cretaceous), Santa Cruz Province, Argentina. Cretaceous Research 154:105735:1-33 (https://doi.org/10.1016/j.cretres.2023.105735)
↑¹K. J. Lacovara, M. C. Lamanna, and L. M. Ibiricu, J. C. Poole, E. R. Schroeter, P. V. Ullmann, K. K. Voegele, Z. M. Boles, A. M. Carter, E. K. Fowler, V. M. Egerton, A. E. Moyer, C. L. Coughenour, J. P. Schein, J. D. Harris, R. D. Martínez, F. E. Novas. 2014. A gigantic, exceptionally complete titanosaurian sauropod dinosaur from southern Patagonia, Argentina. Scientific Reports 4:6196:1-9 (https://doi.org/10.1038/srep06196)
↑^{1 2 3}X. Pereda-Suberbiola, H. Asibia, and X. Murelaga, J. J. Elzorza, J. J. Gomez-Alday. 2000. Taphonomy of the Late Cretaceous dinosaur-bearing beds of the Lano Quarry (Iberian Peninsula). Palaeogeography, Palaeoclimatology, Palaeoecology 157:247-275 (https://doi.org/10.1016/s0031-0182(99)00169-8)
↑¹F. Knoll, R. C. Ridgely, and F. Ortega, J. L. Sanz, L. M. Witmer. 2013. Neurocranial osteology and neuroanatomy of a Late Cretaceous titanosaurian sauropod from Spain (Ampelosaurus sp.). PLoS ONE 8(1):e54991:1-11 (https://doi.org/10.1371/journal.pone.0054991)
↑¹V. Díez Díaz, P. Mocho, and A. Páramo, F. Escaso, F. Marcos-Fernández, J. L. Sanz, F. Ortega. 2016. A new titanosaur (Dinosauria, Sauropoda) from the Upper Cretaceous of Lo Hueco (Cuenca, Spain). Cretaceous Research 68:49-60 (https://doi.org/10.1016/j.cretres.2016.08.001)
↑¹F. Ortega and A. Pérez García. 2009. cf. Lirainosaurus sp. (Dinosauria: Titanosauria) en el Cretácico Superior de Sacedón (Guadalajara). Geogaceta 46:87-90
↑^{1 2 3 4}J. Company, X. Pereda Suberbiola, and J. I. Ruiz-Omenaca. 2009. Nuevos restos fosiles del dinosaurio Lirainosaurus (Sauropoda, Titanosauria) en el Cretácico Superior (Campaniano-Maastrichtiano) de la Península Ibérica. Ameghiniana 46(2):391-405
↑¹J. Le Loeuff. 1995. Ampelosaurus atacis (nov, gen., nov. sp.), un nouveau Titanosauridae (Dinosauria, Sauropoda) du Crétacé supérieur de la Haute Vallée de l'Aude (France) [Ampelosaurus atacis (nov, gen., nov. sp.), a new titanosaurid (Dinosauria, Sauropoda) from the Upper Cretaceous of the Upper Aude Valley (France)]. Comptes Rendus de l'Académie des Sciences à Paris, Série IIa 321:693-699
↑^{1 2 3 4}B. Vila, A. Galobart, and J. I. Canudo, J. Le Loeuff, J. Dinarès-Turell, V. Riera, O. Oms, T. Tortosa, R. Gaete. 2012. The diversity of sauropod dinosaurs and their first taxonomic succession from the latest Cretaceous of southwestern Europe: Clues to demise and extinction. Palaeogeography, Palaeoclimatology, Palaeoecology 350–352:19-38 (https://doi.org/10.1016/j.palaeo.2012.06.008)
↑¹E. Buffetaut. 1998. First evidence of enantiornithine birds from the Upper Cretaceous of Europe: postcranial bones from Cruzy (Hérault, southern France). Oryctos 1:131-136
↑¹V. Díez Díaz, G. Garcia, and X. Pereda Suberbiola, B. Jentgen-Ceschino, K. Stein, P. Godefroit, X. Valentin. 2021. A new titanosaur (Dinosauria: Sauropoda) from the Upper Cretaceous of Velaux-La-Bastide Neuve (southern France). Historical Biology 33(11):2998-3017 (https://doi.org/10.1080/08912963.2020.1841184)
↑¹G. Garcia, S. Amico, and F. Fournier, E. Thouand, X. Valentin. 2010. A new titanosaur genus (Dinosauria, Sauropoda) from the Late Cretaceous of southern France and its paleobiogeographic implications. Bulletin de la Société géologique de France 181(3):269-277 (https://doi.org/10.2113/gssgfbull.181.3.269)
↑^{1 2}J. Le Loeuff, S. Suteethorn, and E. Buffetaut. 2013. A new sauropod dinosaur from the Albian of Le Havre (Normandy, France). Oryctos 10:23-30
↑^{1 2 3 4}S. Brusatte, M. Vremir, and A. Watanabe, Z. Csiki-Sava, D. Naish, G. J. Dyke, G. M. Erickson, M. A. Norell. 2013. An infant ornithopod dinosaur tibia from the Late Cretaceous of Sebes, Romania. Terra Sebus. Acta Musei Sabesiensis 5:627-644
↑¹M. Vremir. 2010. New faunal elements from the Late Cretaceous (Maastrichtian) continental deposits of Sebeş area (Transylvania). Terra Sebus. Acta Musei Sabesiensis 2010(2):635-684
↑¹M. Delfino, V. Codrea, and A. Folie, P. Dica, P. Godefroit, T. Smith. 2008. A complete skull of Allodaposuchus precedens Nopcsa, 1928 (Eusuchia) and a reassessment of the morphology of the taxon based on the Romanian remains. Journal of Vertebrate Paleontology 28(1):111-122 (https://doi.org/10.1671/0272-4634(2008)28[111:acsoap]2.0.co;2)
↑^{1 2 3}V. Díez Díaz, P. D. Mannion, and Z. Csiki-Sava, P. Upchurch. 2025. Revision of Romanian sauropod dinosaurs reveals high titanosaur diversity and body-size disparity on the latest Cretaceous Haţeg Island, with implications for titanosaurian biogeography. Journal of Systematic Palaeontology 23(1):2441516 (https://doi.org/10.1080/14772019.2024.2441516)
↑^{1 2}D. Grigorescu, M. Venczel, and Z. Csiki, R. Limberea. 1999. New latest Cretaceous microvertebrate fossil assemblages from the Haţeg Basin (Romania). Geologie en Mijnbouw 78:301-314
↑¹B. F. Nopcsa. 1915. Die dinosaurier der Siebenbürgischen landesteile Ungarns [Dinosaurs of the Transylvanian regions of Hungary]. Mitteilungen aus den Jahrbuch der Königlich Ungarnischen Geologischen Reichsanstalt 23:1-24
↑¹Z. Csiki-Sava, M. Vremir, and S. Vasile, S. L. Brusatte, G. Dyke, D. Naish, M. A. Norell, R. Totoianu. 2016. The East Side Story – The Transylvanian latest Cretaceous continental vertebrate record and its implications for understanding Cretaceous–Paleogene boundary events. Cretaceous Research 57:662-698 (https://doi.org/10.1016/j.cretres.2015.09.003)
↑¹Z. Csiki, V. Codrea, and C. Jipa-Murzea, P. Godefroit. 2010. A partial titanosaur (Sauropoda, Dinosauria) skeleton from the Maastrichtian of Nalat-Vad, Hateg Basin, Romania. Neues Jahrbuch für Geologie und Paläontologie Abhandlungen 258(3):297-324 (https://doi.org/10.1127/0077-7749/2010/0098)
↑^{1 2}F. v. Huene. 1932. Die fossile Reptil-Ordnung Saurischia, ihre Entwicklung und Geschichte [The fossil reptile order Saurischia, their development and history]. Monographien zur Geologie und Palaeontologie, serie 1 4(1-2):1-361

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