Original description
L. K. Gabuniya and V. V. Kurbatov. 1988. O yurskikh sledakh dinozavrov na yuge sredney Azii [Dinosaur traces in Jurassic deposits of southern Central Asia]. Sledy Zhiznedeyatel'nosti i Dinamika Sredy v Drevnikh Biotopakh. Trudy XXX Sessii Vsesoyuznogo Paleontologicheskogo Obshchestva i VII Sessii Ukrainskogo Paleontologicheskogo Obshchestva [Fossil Traces of Vital Activity and Dynamics of the Environment in Ancient Biotopes. Transactions of the XXX Session of All-Union Paleontological Society and the VII Session of the Ukrainian Paleontological Society]. Naukova Dumka, Kiev