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Ornithopoda

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‘Koutalisaurus kohlerorum’ (Ornithopoda, Lambeosaurinae; indeterminate lambeosaurine sensu Prieto-Márquez et al. 2013), right dentary (IPS 29920 (formerly IPS SRA 27) in medial view (‘lower red unit’ of the Tremp Formation, Les Llaus near Sant Romà d’Abella, Lleida)
Taxons Koutalisaurus

‘Koutalisaurus kohlerorum’ (Ornithopoda, Lambeosaurinae; indeterminate lambeosaurine sensu Prieto-Márquez et al. 2013), right dentary (IPS 29920 (formerly IPS SRA 27) in medial view (‘lower red unit’ of the Tremp Formation, Les Llaus near Sant Romà d’Abella, Lleida)

Koutalisaurus Lambeosaurinae Ornithopoda formation
Theiophytalia kerri Brill & Carpenter, 2006 - ornithopod dinosaur skull from the Cretaceous of Colorado, USA. (YPM 1887, public display, Garden of the Gods visitor center, Colorado Springs, Colorado, USA) This skull is the holotype and only known specimen of an entire genus of ornithopod dinosaurs, Theiophytalia. Ornithopods were herbivorous dinosaurs. From exhibit signage: A Brand New Dinosaur Species Theiophytalia kerri What we know about the dinosaur fossil - so far: It is the only Theiophytalia kerri fossil known to exist in the world. It is a brand new genus and species of dinosaur. Theiophytalia kerri means "belonging to the Garden of the Gods". "Theios" is a Greek word that means "belonging to the gods" and "phytalia" means "garden". "kerri" honors James Hutchinson Kerr (pronounced "Care"), who discovered the dinosaur fossil. Theiophytalia was a medium-sized dinosaur, measuring about 30 feet from head to tail. It was found in 1878 in the Garden of the Gods in the lower Lytle Member of the Purgatoire Rock Formation of the Dakota Group. The dinosaur fossil skull is from the Aptian-Albian Age of the Cretaceous Ear, 125 - 100 million years old. Look closely at the fossil skull. Where are the teeth located? You can see the teeth in the back of the jaw. There are no teeth in the front of the dinosaur's beak-like mouth. This is evidence that the Theiophytalia was a plant eater. It could nip and tear plants, then grind them between its tightly-packed back teeth, similar to plant-eaters of today. Timeline: Dinosaur Fossil Discovered, then Forgotten 1878 - James Kerr, geology professor at Colorado College, finds a fossil skull "in one of the ridges east of the red rocks of the Garden of the Gods." 1886 - O.C. Marsh, famous 1800s dinosaur collector from Yale University, obtains the fossil skull from James Kerr, identifies it as a Camptosaurus dinosaur, and sends the fossil to the Yale Peabody Museum in New Haven, Connecticut. 1886 - 1995 - While the dinosaur fossil skull safely rests in the Yale Museum for many decades, knowledge of its existence is forgotten in Colorado Springs. Rediscovery - 117 years later 1994 - Colorado Springs City park staff research new exhibits for the Garden of the Gods Visitor and Nature Center, set to open in 1995. They meet with Doctor Kirk Johnson, curator of paleontology at the Denver Museum of Nature & Science. 1995 - Kirk Johnson refers the park staff to his museum colleague Doctor Ken Carpenter, expert dinosaur scientist. Doctor Carpenter remembers seeing in his files "something about a dinosaur fossil found in Garden of the Gods" and mails the following article to park staff. Camptosaurus amplus No. 1887, Yale Museum, consisting of portions of the skull and lower jaw. It was collected from deposits in the Garden of the Gods, Colorado Springs, Colorado. With this specimen was found the following note in Professor O.C. Marsh's handwriting, "Part of this animal and various Sauropoda bones were taken out by Professor Kerr in 1878." A Case of Mistaken Identity Reveals a Brand New Dinosaur! 1996 - Kirk Johnson (a Yale alumnus) secures permission to hand-carry the Camptosaurus fossil from the Yale Peabody Museum to Denver so that Ken Carpenter can make a cast (a precise replica) of the fossil. Doctor Carpenter notices irregularities in the Camptosaurus fossil and decides to re-examine the fossil when his schedule permits. 1997 - The Camptosaurus fossil replica is given to the City of Colorado Springs and is exhibited at the Garden of the Gods Visitor and Nature Center. 2006 - Doctor Carpenter and his associate Kathleen Brill reassess the fossil skull and note that it differs from other Camptosaurus skulls in several significant ways, such as the narrower mouth and snout, and the position of the nasal openings, and the bony structures over the eyes. Also, microscopic identification of the rock matrix clinging to the fossil, and research of archival maps, reveal that the fossil skull was actually found in the lower Dakota Rock Formation, not the Morrison Formation as originally reported in the 1800s. The skull is from a dinosaur that lived in the Cretaceous Ear and can't be a Jurassic Era Camptosaurus. Doctor Carpenter's exacting research reveals that the dinosaur skull is a brand new genus and species of dinosaur! He names it Theiophytalia kerri. 2008 - The Theiophytalia kerri is proudly re-exhibited at the Garden of the Gods Visitor and Nature Center. Classification: Animalia, Chordata, Vertebrata, Dinosauria, Ornithischia, Ornithopoda Stratigraphy: lower Lytle Member, Purgatoire Formation, Dakota Group, Aptian to Albian Stages, upper Lower Cretaceous Locality: Garden of the Gods, Colorado Springs, Colorado, USA See info. at: en.wikipedia.org/wiki/Theiophytalia and en.wikipedia.org/wiki/Ornithopoda
Taxons Theiophytalia

Theiophytalia kerri Brill & Carpenter, 2006 - ornithopod dinosaur skull from the Cretaceous of Colorado, USA. (YPM 1887, public display, Garden of the Gods visitor center, Colorado Springs, Colorado, USA) This skull is the holotype and only known specimen of an entire genus of ornithopod dinosaurs, Theiophytalia. Ornithopods were herbivorous dinosaurs. From exhibit signage: A Brand New Dinosaur Species Theiophytalia kerri What we know about the dinosaur fossil - so far: It is the only Theiophytalia kerri fossil known to exist in the world. It is a brand new genus and species of dinosaur. Theiophytalia kerri means "belonging to the Garden of the Gods". "Theios" is a Greek word that means "belonging to the gods" and "phytalia" means "garden". "kerri" honors James Hutchinson Kerr (pronounced "Care"), who discovered the dinosaur fossil. Theiophytalia was a medium-sized dinosaur, measuring about 30 feet from head to tail. It was found in 1878 in the Garden of the Gods in the lower Lytle Member of the Purgatoire Rock Formation of the Dakota Group. The dinosaur fossil skull is from the Aptian-Albian Age of the Cretaceous Ear, 125 - 100 million years old. Look closely at the fossil skull. Where are the teeth located? You can see the teeth in the back of the jaw. There are no teeth in the front of the dinosaur's beak-like mouth. This is evidence that the Theiophytalia was a plant eater. It could nip and tear plants, then grind them between its tightly-packed back teeth, similar to plant-eaters of today. Timeline: Dinosaur Fossil Discovered, then Forgotten 1878 - James Kerr, geology professor at Colorado College, finds a fossil skull "in one of the ridges east of the red rocks of the Garden of the Gods." 1886 - O.C. Marsh, famous 1800s dinosaur collector from Yale University, obtains the fossil skull from James Kerr, identifies it as a Camptosaurus dinosaur, and sends the fossil to the Yale Peabody Museum in New Haven, Connecticut. 1886 - 1995 - While the dinosaur fossil skull safely rests in the Yale Museum for many decades, knowledge of its existence is forgotten in Colorado Springs. Rediscovery - 117 years later 1994 - Colorado Springs City park staff research new exhibits for the Garden of the Gods Visitor and Nature Center, set to open in 1995. They meet with Doctor Kirk Johnson, curator of paleontology at the Denver Museum of Nature & Science. 1995 - Kirk Johnson refers the park staff to his museum colleague Doctor Ken Carpenter, expert dinosaur scientist. Doctor Carpenter remembers seeing in his files "something about a dinosaur fossil found in Garden of the Gods" and mails the following article to park staff. Camptosaurus amplus No. 1887, Yale Museum, consisting of portions of the skull and lower jaw. It was collected from deposits in the Garden of the Gods, Colorado Springs, Colorado. With this specimen was found the following note in Professor O.C. Marsh's handwriting, "Part of this animal and various Sauropoda bones were taken out by Professor Kerr in 1878." A Case of Mistaken Identity Reveals a Brand New Dinosaur! 1996 - Kirk Johnson (a Yale alumnus) secures permission to hand-carry the Camptosaurus fossil from the Yale Peabody Museum to Denver so that Ken Carpenter can make a cast (a precise replica) of the fossil. Doctor Carpenter notices irregularities in the Camptosaurus fossil and decides to re-examine the fossil when his schedule permits. 1997 - The Camptosaurus fossil replica is given to the City of Colorado Springs and is exhibited at the Garden of the Gods Visitor and Nature Center. 2006 - Doctor Carpenter and his associate Kathleen Brill reassess the fossil skull and note that it differs from other Camptosaurus skulls in several significant ways, such as the narrower mouth and snout, and the position of the nasal openings, and the bony structures over the eyes. Also, microscopic identification of the rock matrix clinging to the fossil, and research of archival maps, reveal that the fossil skull was actually found in the lower Dakota Rock Formation, not the Morrison Formation as originally reported in the 1800s. The skull is from a dinosaur that lived in the Cretaceous Ear and can't be a Jurassic Era Camptosaurus. Doctor Carpenter's exacting research reveals that the dinosaur skull is a brand new genus and species of dinosaur! He names it Theiophytalia kerri. 2008 - The Theiophytalia kerri is proudly re-exhibited at the Garden of the Gods Visitor and Nature Center. Classification: Animalia, Chordata, Vertebrata, Dinosauria, Ornithischia, Ornithopoda Stratigraphy: lower Lytle Member, Purgatoire Formation, Dakota Group, Aptian to Albian Stages, upper Lower Cretaceous Locality: Garden of the Gods, Colorado Springs, Colorado, USA See info. at: en.wikipedia.org/wiki/Theiophytalia and en.wikipedia.org/wiki/Ornithopoda

os musée États-Unis Denver +19
Left femur of cf. Iguanodontidae gen. et sp. indet. (Ornithischia, Ornithopoda), IGP MZHLZ/2003/1, in posterior (A), lateral (B), medial (C), ante− rior (D), and distal (F) views, with transverse cross−section through the femoral shaft (E).
Taxons Burianosaurus

Left femur of cf. Iguanodontidae gen. et sp. indet. (Ornithischia, Ornithopoda), IGP MZHLZ/2003/1, in posterior (A), lateral (B), medial (C), ante− rior (D), and distal (F) views, with transverse cross−section through the femoral shaft (E).

Burianosaurus Iguanodontidae Ornithischia Ornithopoda
Seven ornithopods (top to bottom): Ouranosaurus, Tenontosaurus (background) with Convolosaurus (foreground), Muttaburrasaurus, Edmontosaurus annectens, Dryosaurus altus, Corythosaurus casuarius
Taxons Ornithopoda

Seven ornithopods (top to bottom): Ouranosaurus, Tenontosaurus (background) with Convolosaurus (foreground), Muttaburrasaurus, Edmontosaurus annectens, Dryosaurus altus, Corythosaurus casuarius

Convolosaurus Corythosaurus Dryosaurus Edmontosaurus +4
Diagram depicting the currently named Dinosauria from the Late Cretaceous Allen Formation of Argentina. Dinosaur taxa: Aeolosaurus sp. → Salitral Moreno locality, Río Negro Province.[1] Length = 15 meters.[2] Austroraptor cabazai → Santa Rosa Basin locality, Río Negro Province.[3][4] Length = 6 meters.[2] Bonapartenykus ultimus → Salitral Ojo de Agua locality, Río Negro Province.[5] Length = 2.5 meters.[5] Bonapartesaurus rionegrensis → Salitral Moreno locality, Río Negro Province.[6] Length = ∼6 meters.[1] Bonatitan reigi → Salitral de Santa Rosa locality, Río Negro Province.[7] Length = Extrapolated after relatives. Kelumapusaura machi → Cerro Matadero locality, Río Negro Province.[8] 9 meters.[8] Lamarqueavis australis → Cerro Tortugas locality, Río Negro Province.[9] Length = ∼House sparrow-sized.[9] Lapampasaurus cholinoi → Islas Malvinas locality, La Pampa Province.[10] Length = ∼7 meters.[2] Limenavis patagonica → Salitral Moreno locality, Río Negro Province.[11] Length = Extrapolated after relatives. Menucocelsior arriagadai → Salitral Ojo de Agua locality, Río Negro Province. Niebla antiqua → Cerro Matadero locality, Río Negro Province.[12] Length = 4.5 meters.[12] Panamericansaurus schroederi → Bodega Familia Schroeder locality, Neuquén Province.[13] Length = 11 meters.[2] Patagopelta cristata → Salitral Moreno locality, Río Negro Province. Quilmesaurus curriei → Salitral Ojo de Agua locality, Río Negro Province.[14] Length = 5.3 meters.[12] Rocasaurus muniozi → Salitral Moreno locality, Río Negro Province.[15] Length = Extrapolated after relatives. Excluded taxa: Willinakaqe salitralensis is considered a nomen dubium,[16] and its paratype has been reassigned to Bonapartesaurus rionegrensis.[6] Laplatasaurus araukanicus has been restricted to its lectotype which hails from the Anacleto Formation.[17] Abelisaurus comahuensis could either belong to the Allen or Anacleto formations.[12] Pellegrinisaurus powelli could either belong to the Allen or Anacleto formations.[18][19] References ↑ (2013). "The titanosaur sauropods from the late Campanian-early Maastrichtian Allen Formation of Salitral Moreno, Río Negro, Argentina". Acta Palaeontologica Polonica 58 (2): 269–284. DOI:10.4202/app.2011.0055. ↑ a b c (2007) Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages, Random House ISBN: 9780375824197. Genus List for Holtz 2012 Weight Information ↑ (2008). "A bizarre Cretaceous theropod dinosaur from Patagonia and the evolution of Gondwanan dromaeosaurids". Proceedings of the Royal Society B: Biological Sciences 276 (1659): 1101–7. DOI:10.1098/rspb.2008.1554. ISSN 1471-2954. ↑ (2012). "A New Specimen of Austroraptor cabazai Novas, Pol, Canale, Porfiri and Calvo, 2008 (Dinosauria, Theropoda, Unenlagiidae) from the Latest Cretaceous (Maastrichtian) of Río Negro, Argentina". Ameghiniana 49 (4): 662–667. DOI:10.5710/AMGH.30.8.2012.574. ↑ a b Federico L. Agnolin (2012). "New alvarezsaurid (Dinosauria, Theropoda) from uppermost Cretaceous of north-western Patagonia with associated eggs". Cretaceous Research 35: 33–56. DOI:10.1016/j.cretres.2011.11.014. ↑ a b (2017). "Bonapartesaurus rionegrensis, a new hadrosaurine dinosaur from South America: implications for phylogenetic and biogeographic relations with North America". Journal of Vertebrate Paleontology 37 (2): 1–16. DOI:10.1080/02724634.2017.1289381. ↑ Salgado L., Gallina P.A. and Paulina Carabajal A. 2014. "Redescription of Bonatitan reigi (Sauropoda: Titanosauria), from the Campanian–Maastrichtian of the Río Negro Province (Argentina)". Historical Biology: An International Journal of Paleobiology 27(5): 525-548 ↑ a b (2022). "A new hadrosaurid (Dinosauria: Ornithischia) from the Late Cretaceous of northern Patagonia and the radiation of South American hadrosaurids". Journal of Systematic Palaeontology. DOI:10.1080/14772019.2021.2020917. ↑ a b (2010). "[https://pdfs.semanticscholar.org/f6f4/c6eb05d224719916c0b20634f54dfeb37d3f.pdf An avian coracoid from the Upper Cretaceous of Patagonia, Argentina]". Studia Geologica Salmanticensia 46 (2): 99-119. ISSN 0211-8327. ↑ Rodolfo A. Coria, Bernardo González Riga and Silvio Casadío (2012). "Un nuevo hadrosáurido (Dinosauria, Ornithopoda) de la Formación Allen, provincia de La Pampa, Argentina". Ameghiniana 49 (4): 552–572. ↑ Clarke and Chiappe, 2001. A new carinate bird from the Late Cretaceous of Patagonia (Argentina). American Museum Novitates. 3323, 1-23. ↑ a b c d (in English) Aranciaga Rolando, Mauro (2020). "A new medium-sized abelisaurid (Theropoda, Dinosauria) from the late cretaceous (Maastrichtian) Allen Formation of Northern Patagonia, Argentina". Journal of South American Earth Sciences: 102915. DOI:10.1016/j.jsames.2020.102915. ISSN 0895-9811. ↑ (2010). "Panamericansaurus schroederi gen. nov. sp. nov. Un nuevo Sauropoda (Titanosauridae-Aeolosaurini) de la Provincia del Neuquén, Cretácico Superior de Patagonia, Argentina". Brazilian Geographical Journal: Geosciences and Humanities research medium 1: 100–115. ↑ Coria, R.A. (2001) "A new theropod from the Late Cretaceous of Patagonia" in Tanke, Darren H., ed. Mesozoic Vertebrate Life, Life of the Past, Indiana University Press, pp. 3–9 ISBN: 978-0-253-33907-2. ↑ Salgado, L. and C. Azpilicueta. 2000. Un nuevo saltasaurino (Sauropoda, Titanosauridae) de la provincia de Río Negro (Formacíon Allen, Cretácico Superior), Patagonia, Argentina archive copy at the Wayback Machine. Ameghiniana 37 (3):259-264. ↑ (2016). "Revisiting the hadrosaurid diversity of the Allen Fm.: Re-evaluation of the taxonomic validity of Willinakaqe salitralensis (Ornithopoda, Hadrosauridae) from Salitral Moreno, Río Negro Province, Argentina". Ameghiniana 53 (2): 231–237. DOI:10.5710/AMGH.25.09.2015.2943. ↑ Pablo A. Gallina & Alejandro Otero (2015) Reassessment of Laplatasaurus araukanicus (SAUROPODA: TITANOSAURIA), from the Late Cretaceous of Patagonia, Argentina. Ameghiniana 52 (5):487–501. doi:10.5710/AMGH.08.06.2015.2911. ↑ (1996). "Pellegrinisaurus powelli nov. gen. et sp. (Sauropoda, Titanosauridae) from the Upper Cretaceous of Lago Pellegrini, Northwestern Patagonia, Argentina". Ameghiniana 33 (4): 355–365. ISSN 1851-8044. ↑ Heredia, S., & Salgado, L. (2014). Posición estratigráfica de los estratos supracretácicos portadores de dinosaurios en Lago Pellegrini, Patagonia septentrional, Argentina. Ameghiniana, 36(2), 229-234.
Taxons Menucocelsior

Diagram depicting the currently named Dinosauria from the Late Cretaceous Allen Formation of Argentina. Dinosaur taxa: Aeolosaurus sp. → Salitral Moreno locality, Río Negro Province.[1] Length = 15 meters.[2] Austroraptor cabazai → Santa Rosa Basin locality, Río Negro Province.[3][4] Length = 6 meters.[2] Bonapartenykus ultimus → Salitral Ojo de Agua locality, Río Negro Province.[5] Length = 2.5 meters.[5] Bonapartesaurus rionegrensis → Salitral Moreno locality, Río Negro Province.[6] Length = ∼6 meters.[1] Bonatitan reigi → Salitral de Santa Rosa locality, Río Negro Province.[7] Length = Extrapolated after relatives. Kelumapusaura machi → Cerro Matadero locality, Río Negro Province.[8] 9 meters.[8] Lamarqueavis australis → Cerro Tortugas locality, Río Negro Province.[9] Length = ∼House sparrow-sized.[9] Lapampasaurus cholinoi → Islas Malvinas locality, La Pampa Province.[10] Length = ∼7 meters.[2] Limenavis patagonica → Salitral Moreno locality, Río Negro Province.[11] Length = Extrapolated after relatives. Menucocelsior arriagadai → Salitral Ojo de Agua locality, Río Negro Province. Niebla antiqua → Cerro Matadero locality, Río Negro Province.[12] Length = 4.5 meters.[12] Panamericansaurus schroederi → Bodega Familia Schroeder locality, Neuquén Province.[13] Length = 11 meters.[2] Patagopelta cristata → Salitral Moreno locality, Río Negro Province. Quilmesaurus curriei → Salitral Ojo de Agua locality, Río Negro Province.[14] Length = 5.3 meters.[12] Rocasaurus muniozi → Salitral Moreno locality, Río Negro Province.[15] Length = Extrapolated after relatives. Excluded taxa: Willinakaqe salitralensis is considered a nomen dubium,[16] and its paratype has been reassigned to Bonapartesaurus rionegrensis.[6] Laplatasaurus araukanicus has been restricted to its lectotype which hails from the Anacleto Formation.[17] Abelisaurus comahuensis could either belong to the Allen or Anacleto formations.[12] Pellegrinisaurus powelli could either belong to the Allen or Anacleto formations.[18][19] References ↑ (2013). "The titanosaur sauropods from the late Campanian-early Maastrichtian Allen Formation of Salitral Moreno, Río Negro, Argentina". Acta Palaeontologica Polonica 58 (2): 269–284. DOI:10.4202/app.2011.0055. ↑ a b c (2007) Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages, Random House ISBN: 9780375824197. Genus List for Holtz 2012 Weight Information ↑ (2008). "A bizarre Cretaceous theropod dinosaur from Patagonia and the evolution of Gondwanan dromaeosaurids". Proceedings of the Royal Society B: Biological Sciences 276 (1659): 1101–7. DOI:10.1098/rspb.2008.1554. ISSN 1471-2954. ↑ (2012). "A New Specimen of Austroraptor cabazai Novas, Pol, Canale, Porfiri and Calvo, 2008 (Dinosauria, Theropoda, Unenlagiidae) from the Latest Cretaceous (Maastrichtian) of Río Negro, Argentina". Ameghiniana 49 (4): 662–667. DOI:10.5710/AMGH.30.8.2012.574. ↑ a b Federico L. Agnolin (2012). "New alvarezsaurid (Dinosauria, Theropoda) from uppermost Cretaceous of north-western Patagonia with associated eggs". Cretaceous Research 35: 33–56. DOI:10.1016/j.cretres.2011.11.014. ↑ a b (2017). "Bonapartesaurus rionegrensis, a new hadrosaurine dinosaur from South America: implications for phylogenetic and biogeographic relations with North America". Journal of Vertebrate Paleontology 37 (2): 1–16. DOI:10.1080/02724634.2017.1289381. ↑ Salgado L., Gallina P.A. and Paulina Carabajal A. 2014. "Redescription of Bonatitan reigi (Sauropoda: Titanosauria), from the Campanian–Maastrichtian of the Río Negro Province (Argentina)". Historical Biology: An International Journal of Paleobiology 27(5): 525-548 ↑ a b (2022). "A new hadrosaurid (Dinosauria: Ornithischia) from the Late Cretaceous of northern Patagonia and the radiation of South American hadrosaurids". Journal of Systematic Palaeontology. DOI:10.1080/14772019.2021.2020917. ↑ a b (2010). "[https://pdfs.semanticscholar.org/f6f4/c6eb05d224719916c0b20634f54dfeb37d3f.pdf An avian coracoid from the Upper Cretaceous of Patagonia, Argentina]". Studia Geologica Salmanticensia 46 (2): 99-119. ISSN 0211-8327. ↑ Rodolfo A. Coria, Bernardo González Riga and Silvio Casadío (2012). "Un nuevo hadrosáurido (Dinosauria, Ornithopoda) de la Formación Allen, provincia de La Pampa, Argentina". Ameghiniana 49 (4): 552–572. ↑ Clarke and Chiappe, 2001. A new carinate bird from the Late Cretaceous of Patagonia (Argentina). American Museum Novitates. 3323, 1-23. ↑ a b c d (in English) Aranciaga Rolando, Mauro (2020). "A new medium-sized abelisaurid (Theropoda, Dinosauria) from the late cretaceous (Maastrichtian) Allen Formation of Northern Patagonia, Argentina". Journal of South American Earth Sciences: 102915. DOI:10.1016/j.jsames.2020.102915. ISSN 0895-9811. ↑ (2010). "Panamericansaurus schroederi gen. nov. sp. nov. Un nuevo Sauropoda (Titanosauridae-Aeolosaurini) de la Provincia del Neuquén, Cretácico Superior de Patagonia, Argentina". Brazilian Geographical Journal: Geosciences and Humanities research medium 1: 100–115. ↑ Coria, R.A. (2001) "A new theropod from the Late Cretaceous of Patagonia" in Tanke, Darren H., ed. Mesozoic Vertebrate Life, Life of the Past, Indiana University Press, pp. 3–9 ISBN: 978-0-253-33907-2. ↑ Salgado, L. and C. Azpilicueta. 2000. Un nuevo saltasaurino (Sauropoda, Titanosauridae) de la provincia de Río Negro (Formacíon Allen, Cretácico Superior), Patagonia, Argentina archive copy at the Wayback Machine. Ameghiniana 37 (3):259-264. ↑ (2016). "Revisiting the hadrosaurid diversity of the Allen Fm.: Re-evaluation of the taxonomic validity of Willinakaqe salitralensis (Ornithopoda, Hadrosauridae) from Salitral Moreno, Río Negro Province, Argentina". Ameghiniana 53 (2): 231–237. DOI:10.5710/AMGH.25.09.2015.2943. ↑ Pablo A. Gallina & Alejandro Otero (2015) Reassessment of Laplatasaurus araukanicus (SAUROPODA: TITANOSAURIA), from the Late Cretaceous of Patagonia, Argentina. Ameghiniana 52 (5):487–501. doi:10.5710/AMGH.08.06.2015.2911. ↑ (1996). "Pellegrinisaurus powelli nov. gen. et sp. (Sauropoda, Titanosauridae) from the Upper Cretaceous of Lago Pellegrini, Northwestern Patagonia, Argentina". Ameghiniana 33 (4): 355–365. ISSN 1851-8044. ↑ Heredia, S., & Salgado, L. (2014). Posición estratigráfica de los estratos supracretácicos portadores de dinosaurios en Lago Pellegrini, Patagonia septentrional, Argentina. Ameghiniana, 36(2), 229-234.

musée Argentine Allen Anacleto +39
Diagram depicting the currently named Dinosauria from the Late Cretaceous Allen Formation of Argentina. Dinosaur taxa: Aeolosaurus sp. → Salitral Moreno locality, Río Negro Province.[1] Length = 15 meters.[2] Austroraptor cabazai → Santa Rosa Basin locality, Río Negro Province.[3][4] Length = 6 meters.[2] Bonapartenykus ultimus → Salitral Ojo de Agua locality, Río Negro Province.[5] Length = 2.5 meters.[5] Bonapartesaurus rionegrensis → Salitral Moreno locality, Río Negro Province.[6] Length = ∼6 meters.[1] Bonatitan reigi → Salitral de Santa Rosa locality, Río Negro Province.[7] Length = Extrapolated after relatives. Kelumapusaura machi → Cerro Matadero locality, Río Negro Province.[8] 9 meters.[8] Lamarqueavis australis → Cerro Tortugas locality, Río Negro Province.[9] Length = ∼House sparrow-sized.[9] Lapampasaurus cholinoi → Islas Malvinas locality, La Pampa Province.[10] Length = ∼7 meters.[2] Limenavis patagonica → Salitral Moreno locality, Río Negro Province.[11] Length = Extrapolated after relatives. Menucocelsior arriagadai → Salitral Ojo de Agua locality, Río Negro Province. Niebla antiqua → Cerro Matadero locality, Río Negro Province.[12] Length = 4.5 meters.[12] Panamericansaurus schroederi → Bodega Familia Schroeder locality, Neuquén Province.[13] Length = 11 meters.[2] Patagopelta cristata → Salitral Moreno locality, Río Negro Province. Quilmesaurus curriei → Salitral Ojo de Agua locality, Río Negro Province.[14] Length = 5.3 meters.[12] Rocasaurus muniozi → Salitral Moreno locality, Río Negro Province.[15] Length = Extrapolated after relatives. Excluded taxa: Willinakaqe salitralensis is considered a nomen dubium,[16] and its paratype has been reassigned to Bonapartesaurus rionegrensis.[6] Laplatasaurus araukanicus has been restricted to its lectotype which hails from the Anacleto Formation.[17] Abelisaurus comahuensis could either belong to the Allen or Anacleto formations.[12] Pellegrinisaurus powelli could either belong to the Allen or Anacleto formations.[18][19] References ↑ (2013). "The titanosaur sauropods from the late Campanian-early Maastrichtian Allen Formation of Salitral Moreno, Río Negro, Argentina". Acta Palaeontologica Polonica 58 (2): 269–284. DOI:10.4202/app.2011.0055. ↑ a b c (2007) Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages, Random House ISBN: 9780375824197. Genus List for Holtz 2012 Weight Information ↑ (2008). "A bizarre Cretaceous theropod dinosaur from Patagonia and the evolution of Gondwanan dromaeosaurids". Proceedings of the Royal Society B: Biological Sciences 276 (1659): 1101–7. DOI:10.1098/rspb.2008.1554. ISSN 1471-2954. ↑ (2012). "A New Specimen of Austroraptor cabazai Novas, Pol, Canale, Porfiri and Calvo, 2008 (Dinosauria, Theropoda, Unenlagiidae) from the Latest Cretaceous (Maastrichtian) of Río Negro, Argentina". Ameghiniana 49 (4): 662–667. DOI:10.5710/AMGH.30.8.2012.574. ↑ a b Federico L. Agnolin (2012). "New alvarezsaurid (Dinosauria, Theropoda) from uppermost Cretaceous of north-western Patagonia with associated eggs". Cretaceous Research 35: 33–56. DOI:10.1016/j.cretres.2011.11.014. ↑ a b (2017). "Bonapartesaurus rionegrensis, a new hadrosaurine dinosaur from South America: implications for phylogenetic and biogeographic relations with North America". Journal of Vertebrate Paleontology 37 (2): 1–16. DOI:10.1080/02724634.2017.1289381. ↑ Salgado L., Gallina P.A. and Paulina Carabajal A. 2014. "Redescription of Bonatitan reigi (Sauropoda: Titanosauria), from the Campanian–Maastrichtian of the Río Negro Province (Argentina)". Historical Biology: An International Journal of Paleobiology 27(5): 525-548 ↑ a b (2022). "A new hadrosaurid (Dinosauria: Ornithischia) from the Late Cretaceous of northern Patagonia and the radiation of South American hadrosaurids". Journal of Systematic Palaeontology. DOI:10.1080/14772019.2021.2020917. ↑ a b (2010). "[https://pdfs.semanticscholar.org/f6f4/c6eb05d224719916c0b20634f54dfeb37d3f.pdf An avian coracoid from the Upper Cretaceous of Patagonia, Argentina]". Studia Geologica Salmanticensia 46 (2): 99-119. ISSN 0211-8327. ↑ Rodolfo A. Coria, Bernardo González Riga and Silvio Casadío (2012). "Un nuevo hadrosáurido (Dinosauria, Ornithopoda) de la Formación Allen, provincia de La Pampa, Argentina". Ameghiniana 49 (4): 552–572. ↑ Clarke and Chiappe, 2001. A new carinate bird from the Late Cretaceous of Patagonia (Argentina). American Museum Novitates. 3323, 1-23. ↑ a b c d (in English) Aranciaga Rolando, Mauro (2020). "A new medium-sized abelisaurid (Theropoda, Dinosauria) from the late cretaceous (Maastrichtian) Allen Formation of Northern Patagonia, Argentina". Journal of South American Earth Sciences: 102915. DOI:10.1016/j.jsames.2020.102915. ISSN 0895-9811. ↑ (2010). "Panamericansaurus schroederi gen. nov. sp. nov. Un nuevo Sauropoda (Titanosauridae-Aeolosaurini) de la Provincia del Neuquén, Cretácico Superior de Patagonia, Argentina". Brazilian Geographical Journal: Geosciences and Humanities research medium 1: 100–115. ↑ Coria, R.A. (2001) "A new theropod from the Late Cretaceous of Patagonia" in Tanke, Darren H., ed. Mesozoic Vertebrate Life, Life of the Past, Indiana University Press, pp. 3–9 ISBN: 978-0-253-33907-2. ↑ Salgado, L. and C. Azpilicueta. 2000. Un nuevo saltasaurino (Sauropoda, Titanosauridae) de la provincia de Río Negro (Formacíon Allen, Cretácico Superior), Patagonia, Argentina archive copy at the Wayback Machine. Ameghiniana 37 (3):259-264. ↑ (2016). "Revisiting the hadrosaurid diversity of the Allen Fm.: Re-evaluation of the taxonomic validity of Willinakaqe salitralensis (Ornithopoda, Hadrosauridae) from Salitral Moreno, Río Negro Province, Argentina". Ameghiniana 53 (2): 231–237. DOI:10.5710/AMGH.25.09.2015.2943. ↑ Pablo A. Gallina & Alejandro Otero (2015) Reassessment of Laplatasaurus araukanicus (SAUROPODA: TITANOSAURIA), from the Late Cretaceous of Patagonia, Argentina. Ameghiniana 52 (5):487–501. doi:10.5710/AMGH.08.06.2015.2911. ↑ (1996). "Pellegrinisaurus powelli nov. gen. et sp. (Sauropoda, Titanosauridae) from the Upper Cretaceous of Lago Pellegrini, Northwestern Patagonia, Argentina". Ameghiniana 33 (4): 355–365. ISSN 1851-8044. ↑ Heredia, S., & Salgado, L. (2014). Posición estratigráfica de los estratos supracretácicos portadores de dinosaurios en Lago Pellegrini, Patagonia septentrional, Argentina. Ameghiniana, 36(2), 229-234.
Taxons Lapampasaurus

Diagram depicting the currently named Dinosauria from the Late Cretaceous Allen Formation of Argentina. Dinosaur taxa: Aeolosaurus sp. → Salitral Moreno locality, Río Negro Province.[1] Length = 15 meters.[2] Austroraptor cabazai → Santa Rosa Basin locality, Río Negro Province.[3][4] Length = 6 meters.[2] Bonapartenykus ultimus → Salitral Ojo de Agua locality, Río Negro Province.[5] Length = 2.5 meters.[5] Bonapartesaurus rionegrensis → Salitral Moreno locality, Río Negro Province.[6] Length = ∼6 meters.[1] Bonatitan reigi → Salitral de Santa Rosa locality, Río Negro Province.[7] Length = Extrapolated after relatives. Kelumapusaura machi → Cerro Matadero locality, Río Negro Province.[8] 9 meters.[8] Lamarqueavis australis → Cerro Tortugas locality, Río Negro Province.[9] Length = ∼House sparrow-sized.[9] Lapampasaurus cholinoi → Islas Malvinas locality, La Pampa Province.[10] Length = ∼7 meters.[2] Limenavis patagonica → Salitral Moreno locality, Río Negro Province.[11] Length = Extrapolated after relatives. Menucocelsior arriagadai → Salitral Ojo de Agua locality, Río Negro Province. Niebla antiqua → Cerro Matadero locality, Río Negro Province.[12] Length = 4.5 meters.[12] Panamericansaurus schroederi → Bodega Familia Schroeder locality, Neuquén Province.[13] Length = 11 meters.[2] Patagopelta cristata → Salitral Moreno locality, Río Negro Province. Quilmesaurus curriei → Salitral Ojo de Agua locality, Río Negro Province.[14] Length = 5.3 meters.[12] Rocasaurus muniozi → Salitral Moreno locality, Río Negro Province.[15] Length = Extrapolated after relatives. Excluded taxa: Willinakaqe salitralensis is considered a nomen dubium,[16] and its paratype has been reassigned to Bonapartesaurus rionegrensis.[6] Laplatasaurus araukanicus has been restricted to its lectotype which hails from the Anacleto Formation.[17] Abelisaurus comahuensis could either belong to the Allen or Anacleto formations.[12] Pellegrinisaurus powelli could either belong to the Allen or Anacleto formations.[18][19] References ↑ (2013). "The titanosaur sauropods from the late Campanian-early Maastrichtian Allen Formation of Salitral Moreno, Río Negro, Argentina". Acta Palaeontologica Polonica 58 (2): 269–284. DOI:10.4202/app.2011.0055. ↑ a b c (2007) Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages, Random House ISBN: 9780375824197. Genus List for Holtz 2012 Weight Information ↑ (2008). "A bizarre Cretaceous theropod dinosaur from Patagonia and the evolution of Gondwanan dromaeosaurids". Proceedings of the Royal Society B: Biological Sciences 276 (1659): 1101–7. DOI:10.1098/rspb.2008.1554. ISSN 1471-2954. ↑ (2012). "A New Specimen of Austroraptor cabazai Novas, Pol, Canale, Porfiri and Calvo, 2008 (Dinosauria, Theropoda, Unenlagiidae) from the Latest Cretaceous (Maastrichtian) of Río Negro, Argentina". Ameghiniana 49 (4): 662–667. DOI:10.5710/AMGH.30.8.2012.574. ↑ a b Federico L. Agnolin (2012). "New alvarezsaurid (Dinosauria, Theropoda) from uppermost Cretaceous of north-western Patagonia with associated eggs". Cretaceous Research 35: 33–56. DOI:10.1016/j.cretres.2011.11.014. ↑ a b (2017). "Bonapartesaurus rionegrensis, a new hadrosaurine dinosaur from South America: implications for phylogenetic and biogeographic relations with North America". Journal of Vertebrate Paleontology 37 (2): 1–16. DOI:10.1080/02724634.2017.1289381. ↑ Salgado L., Gallina P.A. and Paulina Carabajal A. 2014. "Redescription of Bonatitan reigi (Sauropoda: Titanosauria), from the Campanian–Maastrichtian of the Río Negro Province (Argentina)". Historical Biology: An International Journal of Paleobiology 27(5): 525-548 ↑ a b (2022). "A new hadrosaurid (Dinosauria: Ornithischia) from the Late Cretaceous of northern Patagonia and the radiation of South American hadrosaurids". Journal of Systematic Palaeontology. DOI:10.1080/14772019.2021.2020917. ↑ a b (2010). "[https://pdfs.semanticscholar.org/f6f4/c6eb05d224719916c0b20634f54dfeb37d3f.pdf An avian coracoid from the Upper Cretaceous of Patagonia, Argentina]". Studia Geologica Salmanticensia 46 (2): 99-119. ISSN 0211-8327. ↑ Rodolfo A. Coria, Bernardo González Riga and Silvio Casadío (2012). "Un nuevo hadrosáurido (Dinosauria, Ornithopoda) de la Formación Allen, provincia de La Pampa, Argentina". Ameghiniana 49 (4): 552–572. ↑ Clarke and Chiappe, 2001. A new carinate bird from the Late Cretaceous of Patagonia (Argentina). American Museum Novitates. 3323, 1-23. ↑ a b c d (in English) Aranciaga Rolando, Mauro (2020). "A new medium-sized abelisaurid (Theropoda, Dinosauria) from the late cretaceous (Maastrichtian) Allen Formation of Northern Patagonia, Argentina". Journal of South American Earth Sciences: 102915. DOI:10.1016/j.jsames.2020.102915. ISSN 0895-9811. ↑ (2010). "Panamericansaurus schroederi gen. nov. sp. nov. Un nuevo Sauropoda (Titanosauridae-Aeolosaurini) de la Provincia del Neuquén, Cretácico Superior de Patagonia, Argentina". Brazilian Geographical Journal: Geosciences and Humanities research medium 1: 100–115. ↑ Coria, R.A. (2001) "A new theropod from the Late Cretaceous of Patagonia" in Tanke, Darren H., ed. Mesozoic Vertebrate Life, Life of the Past, Indiana University Press, pp. 3–9 ISBN: 978-0-253-33907-2. ↑ Salgado, L. and C. Azpilicueta. 2000. Un nuevo saltasaurino (Sauropoda, Titanosauridae) de la provincia de Río Negro (Formacíon Allen, Cretácico Superior), Patagonia, Argentina archive copy at the Wayback Machine. Ameghiniana 37 (3):259-264. ↑ (2016). "Revisiting the hadrosaurid diversity of the Allen Fm.: Re-evaluation of the taxonomic validity of Willinakaqe salitralensis (Ornithopoda, Hadrosauridae) from Salitral Moreno, Río Negro Province, Argentina". Ameghiniana 53 (2): 231–237. DOI:10.5710/AMGH.25.09.2015.2943. ↑ Pablo A. Gallina & Alejandro Otero (2015) Reassessment of Laplatasaurus araukanicus (SAUROPODA: TITANOSAURIA), from the Late Cretaceous of Patagonia, Argentina. Ameghiniana 52 (5):487–501. doi:10.5710/AMGH.08.06.2015.2911. ↑ (1996). "Pellegrinisaurus powelli nov. gen. et sp. (Sauropoda, Titanosauridae) from the Upper Cretaceous of Lago Pellegrini, Northwestern Patagonia, Argentina". Ameghiniana 33 (4): 355–365. ISSN 1851-8044. ↑ Heredia, S., & Salgado, L. (2014). Posición estratigráfica de los estratos supracretácicos portadores de dinosaurios en Lago Pellegrini, Patagonia septentrional, Argentina. Ameghiniana, 36(2), 229-234.

musée Argentine Allen Anacleto +39
Diagram depicting the currently named Dinosauria from the Late Cretaceous Allen Formation of Argentina. Dinosaur taxa: Aeolosaurus sp. → Salitral Moreno locality, Río Negro Province.[1] Length = 15 meters.[2] Austroraptor cabazai → Santa Rosa Basin locality, Río Negro Province.[3][4] Length = 6 meters.[2] Bonapartenykus ultimus → Salitral Ojo de Agua locality, Río Negro Province.[5] Length = 2.5 meters.[5] Bonapartesaurus rionegrensis → Salitral Moreno locality, Río Negro Province.[6] Length = ∼6 meters.[1] Bonatitan reigi → Salitral de Santa Rosa locality, Río Negro Province.[7] Length = Extrapolated after relatives. Kelumapusaura machi → Cerro Matadero locality, Río Negro Province.[8] 9 meters.[8] Lamarqueavis australis → Cerro Tortugas locality, Río Negro Province.[9] Length = ∼House sparrow-sized.[9] Lapampasaurus cholinoi → Islas Malvinas locality, La Pampa Province.[10] Length = ∼7 meters.[2] Limenavis patagonica → Salitral Moreno locality, Río Negro Province.[11] Length = Extrapolated after relatives. Menucocelsior arriagadai → Salitral Ojo de Agua locality, Río Negro Province. Niebla antiqua → Cerro Matadero locality, Río Negro Province.[12] Length = 4.5 meters.[12] Panamericansaurus schroederi → Bodega Familia Schroeder locality, Neuquén Province.[13] Length = 11 meters.[2] Patagopelta cristata → Salitral Moreno locality, Río Negro Province. Quilmesaurus curriei → Salitral Ojo de Agua locality, Río Negro Province.[14] Length = 5.3 meters.[12] Rocasaurus muniozi → Salitral Moreno locality, Río Negro Province.[15] Length = Extrapolated after relatives. Excluded taxa: Willinakaqe salitralensis is considered a nomen dubium,[16] and its paratype has been reassigned to Bonapartesaurus rionegrensis.[6] Laplatasaurus araukanicus has been restricted to its lectotype which hails from the Anacleto Formation.[17] Abelisaurus comahuensis could either belong to the Allen or Anacleto formations.[12] Pellegrinisaurus powelli could either belong to the Allen or Anacleto formations.[18][19] References ↑ (2013). "The titanosaur sauropods from the late Campanian-early Maastrichtian Allen Formation of Salitral Moreno, Río Negro, Argentina". Acta Palaeontologica Polonica 58 (2): 269–284. DOI:10.4202/app.2011.0055. ↑ a b c (2007) Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages, Random House ISBN: 9780375824197. Genus List for Holtz 2012 Weight Information ↑ (2008). "A bizarre Cretaceous theropod dinosaur from Patagonia and the evolution of Gondwanan dromaeosaurids". Proceedings of the Royal Society B: Biological Sciences 276 (1659): 1101–7. DOI:10.1098/rspb.2008.1554. ISSN 1471-2954. ↑ (2012). "A New Specimen of Austroraptor cabazai Novas, Pol, Canale, Porfiri and Calvo, 2008 (Dinosauria, Theropoda, Unenlagiidae) from the Latest Cretaceous (Maastrichtian) of Río Negro, Argentina". Ameghiniana 49 (4): 662–667. DOI:10.5710/AMGH.30.8.2012.574. ↑ a b Federico L. Agnolin (2012). "New alvarezsaurid (Dinosauria, Theropoda) from uppermost Cretaceous of north-western Patagonia with associated eggs". Cretaceous Research 35: 33–56. DOI:10.1016/j.cretres.2011.11.014. ↑ a b (2017). "Bonapartesaurus rionegrensis, a new hadrosaurine dinosaur from South America: implications for phylogenetic and biogeographic relations with North America". Journal of Vertebrate Paleontology 37 (2): 1–16. DOI:10.1080/02724634.2017.1289381. ↑ Salgado L., Gallina P.A. and Paulina Carabajal A. 2014. "Redescription of Bonatitan reigi (Sauropoda: Titanosauria), from the Campanian–Maastrichtian of the Río Negro Province (Argentina)". Historical Biology: An International Journal of Paleobiology 27(5): 525-548 ↑ a b (2022). "A new hadrosaurid (Dinosauria: Ornithischia) from the Late Cretaceous of northern Patagonia and the radiation of South American hadrosaurids". Journal of Systematic Palaeontology. DOI:10.1080/14772019.2021.2020917. ↑ a b (2010). "[https://pdfs.semanticscholar.org/f6f4/c6eb05d224719916c0b20634f54dfeb37d3f.pdf An avian coracoid from the Upper Cretaceous of Patagonia, Argentina]". Studia Geologica Salmanticensia 46 (2): 99-119. ISSN 0211-8327. ↑ Rodolfo A. Coria, Bernardo González Riga and Silvio Casadío (2012). "Un nuevo hadrosáurido (Dinosauria, Ornithopoda) de la Formación Allen, provincia de La Pampa, Argentina". Ameghiniana 49 (4): 552–572. ↑ Clarke and Chiappe, 2001. A new carinate bird from the Late Cretaceous of Patagonia (Argentina). American Museum Novitates. 3323, 1-23. ↑ a b c d (in English) Aranciaga Rolando, Mauro (2020). "A new medium-sized abelisaurid (Theropoda, Dinosauria) from the late cretaceous (Maastrichtian) Allen Formation of Northern Patagonia, Argentina". Journal of South American Earth Sciences: 102915. DOI:10.1016/j.jsames.2020.102915. ISSN 0895-9811. ↑ (2010). "Panamericansaurus schroederi gen. nov. sp. nov. Un nuevo Sauropoda (Titanosauridae-Aeolosaurini) de la Provincia del Neuquén, Cretácico Superior de Patagonia, Argentina". Brazilian Geographical Journal: Geosciences and Humanities research medium 1: 100–115. ↑ Coria, R.A. (2001) "A new theropod from the Late Cretaceous of Patagonia" in Tanke, Darren H., ed. Mesozoic Vertebrate Life, Life of the Past, Indiana University Press, pp. 3–9 ISBN: 978-0-253-33907-2. ↑ Salgado, L. and C. Azpilicueta. 2000. Un nuevo saltasaurino (Sauropoda, Titanosauridae) de la provincia de Río Negro (Formacíon Allen, Cretácico Superior), Patagonia, Argentina archive copy at the Wayback Machine. Ameghiniana 37 (3):259-264. ↑ (2016). "Revisiting the hadrosaurid diversity of the Allen Fm.: Re-evaluation of the taxonomic validity of Willinakaqe salitralensis (Ornithopoda, Hadrosauridae) from Salitral Moreno, Río Negro Province, Argentina". Ameghiniana 53 (2): 231–237. DOI:10.5710/AMGH.25.09.2015.2943. ↑ Pablo A. Gallina & Alejandro Otero (2015) Reassessment of Laplatasaurus araukanicus (SAUROPODA: TITANOSAURIA), from the Late Cretaceous of Patagonia, Argentina. Ameghiniana 52 (5):487–501. doi:10.5710/AMGH.08.06.2015.2911. ↑ (1996). "Pellegrinisaurus powelli nov. gen. et sp. (Sauropoda, Titanosauridae) from the Upper Cretaceous of Lago Pellegrini, Northwestern Patagonia, Argentina". Ameghiniana 33 (4): 355–365. ISSN 1851-8044. ↑ Heredia, S., & Salgado, L. (2014). Posición estratigráfica de los estratos supracretácicos portadores de dinosaurios en Lago Pellegrini, Patagonia septentrional, Argentina. Ameghiniana, 36(2), 229-234.
Taxons Panamericansaurus

Diagram depicting the currently named Dinosauria from the Late Cretaceous Allen Formation of Argentina. Dinosaur taxa: Aeolosaurus sp. → Salitral Moreno locality, Río Negro Province.[1] Length = 15 meters.[2] Austroraptor cabazai → Santa Rosa Basin locality, Río Negro Province.[3][4] Length = 6 meters.[2] Bonapartenykus ultimus → Salitral Ojo de Agua locality, Río Negro Province.[5] Length = 2.5 meters.[5] Bonapartesaurus rionegrensis → Salitral Moreno locality, Río Negro Province.[6] Length = ∼6 meters.[1] Bonatitan reigi → Salitral de Santa Rosa locality, Río Negro Province.[7] Length = Extrapolated after relatives. Kelumapusaura machi → Cerro Matadero locality, Río Negro Province.[8] 9 meters.[8] Lamarqueavis australis → Cerro Tortugas locality, Río Negro Province.[9] Length = ∼House sparrow-sized.[9] Lapampasaurus cholinoi → Islas Malvinas locality, La Pampa Province.[10] Length = ∼7 meters.[2] Limenavis patagonica → Salitral Moreno locality, Río Negro Province.[11] Length = Extrapolated after relatives. Menucocelsior arriagadai → Salitral Ojo de Agua locality, Río Negro Province. Niebla antiqua → Cerro Matadero locality, Río Negro Province.[12] Length = 4.5 meters.[12] Panamericansaurus schroederi → Bodega Familia Schroeder locality, Neuquén Province.[13] Length = 11 meters.[2] Patagopelta cristata → Salitral Moreno locality, Río Negro Province. Quilmesaurus curriei → Salitral Ojo de Agua locality, Río Negro Province.[14] Length = 5.3 meters.[12] Rocasaurus muniozi → Salitral Moreno locality, Río Negro Province.[15] Length = Extrapolated after relatives. Excluded taxa: Willinakaqe salitralensis is considered a nomen dubium,[16] and its paratype has been reassigned to Bonapartesaurus rionegrensis.[6] Laplatasaurus araukanicus has been restricted to its lectotype which hails from the Anacleto Formation.[17] Abelisaurus comahuensis could either belong to the Allen or Anacleto formations.[12] Pellegrinisaurus powelli could either belong to the Allen or Anacleto formations.[18][19] References ↑ (2013). "The titanosaur sauropods from the late Campanian-early Maastrichtian Allen Formation of Salitral Moreno, Río Negro, Argentina". Acta Palaeontologica Polonica 58 (2): 269–284. DOI:10.4202/app.2011.0055. ↑ a b c (2007) Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages, Random House ISBN: 9780375824197. Genus List for Holtz 2012 Weight Information ↑ (2008). "A bizarre Cretaceous theropod dinosaur from Patagonia and the evolution of Gondwanan dromaeosaurids". Proceedings of the Royal Society B: Biological Sciences 276 (1659): 1101–7. DOI:10.1098/rspb.2008.1554. ISSN 1471-2954. ↑ (2012). "A New Specimen of Austroraptor cabazai Novas, Pol, Canale, Porfiri and Calvo, 2008 (Dinosauria, Theropoda, Unenlagiidae) from the Latest Cretaceous (Maastrichtian) of Río Negro, Argentina". Ameghiniana 49 (4): 662–667. DOI:10.5710/AMGH.30.8.2012.574. ↑ a b Federico L. Agnolin (2012). "New alvarezsaurid (Dinosauria, Theropoda) from uppermost Cretaceous of north-western Patagonia with associated eggs". Cretaceous Research 35: 33–56. DOI:10.1016/j.cretres.2011.11.014. ↑ a b (2017). "Bonapartesaurus rionegrensis, a new hadrosaurine dinosaur from South America: implications for phylogenetic and biogeographic relations with North America". Journal of Vertebrate Paleontology 37 (2): 1–16. DOI:10.1080/02724634.2017.1289381. ↑ Salgado L., Gallina P.A. and Paulina Carabajal A. 2014. "Redescription of Bonatitan reigi (Sauropoda: Titanosauria), from the Campanian–Maastrichtian of the Río Negro Province (Argentina)". Historical Biology: An International Journal of Paleobiology 27(5): 525-548 ↑ a b (2022). "A new hadrosaurid (Dinosauria: Ornithischia) from the Late Cretaceous of northern Patagonia and the radiation of South American hadrosaurids". Journal of Systematic Palaeontology. DOI:10.1080/14772019.2021.2020917. ↑ a b (2010). "[https://pdfs.semanticscholar.org/f6f4/c6eb05d224719916c0b20634f54dfeb37d3f.pdf An avian coracoid from the Upper Cretaceous of Patagonia, Argentina]". Studia Geologica Salmanticensia 46 (2): 99-119. ISSN 0211-8327. ↑ Rodolfo A. Coria, Bernardo González Riga and Silvio Casadío (2012). "Un nuevo hadrosáurido (Dinosauria, Ornithopoda) de la Formación Allen, provincia de La Pampa, Argentina". Ameghiniana 49 (4): 552–572. ↑ Clarke and Chiappe, 2001. A new carinate bird from the Late Cretaceous of Patagonia (Argentina). American Museum Novitates. 3323, 1-23. ↑ a b c d (in English) Aranciaga Rolando, Mauro (2020). "A new medium-sized abelisaurid (Theropoda, Dinosauria) from the late cretaceous (Maastrichtian) Allen Formation of Northern Patagonia, Argentina". Journal of South American Earth Sciences: 102915. DOI:10.1016/j.jsames.2020.102915. ISSN 0895-9811. ↑ (2010). "Panamericansaurus schroederi gen. nov. sp. nov. Un nuevo Sauropoda (Titanosauridae-Aeolosaurini) de la Provincia del Neuquén, Cretácico Superior de Patagonia, Argentina". Brazilian Geographical Journal: Geosciences and Humanities research medium 1: 100–115. ↑ Coria, R.A. (2001) "A new theropod from the Late Cretaceous of Patagonia" in Tanke, Darren H., ed. Mesozoic Vertebrate Life, Life of the Past, Indiana University Press, pp. 3–9 ISBN: 978-0-253-33907-2. ↑ Salgado, L. and C. Azpilicueta. 2000. Un nuevo saltasaurino (Sauropoda, Titanosauridae) de la provincia de Río Negro (Formacíon Allen, Cretácico Superior), Patagonia, Argentina archive copy at the Wayback Machine. Ameghiniana 37 (3):259-264. ↑ (2016). "Revisiting the hadrosaurid diversity of the Allen Fm.: Re-evaluation of the taxonomic validity of Willinakaqe salitralensis (Ornithopoda, Hadrosauridae) from Salitral Moreno, Río Negro Province, Argentina". Ameghiniana 53 (2): 231–237. DOI:10.5710/AMGH.25.09.2015.2943. ↑ Pablo A. Gallina & Alejandro Otero (2015) Reassessment of Laplatasaurus araukanicus (SAUROPODA: TITANOSAURIA), from the Late Cretaceous of Patagonia, Argentina. Ameghiniana 52 (5):487–501. doi:10.5710/AMGH.08.06.2015.2911. ↑ (1996). "Pellegrinisaurus powelli nov. gen. et sp. (Sauropoda, Titanosauridae) from the Upper Cretaceous of Lago Pellegrini, Northwestern Patagonia, Argentina". Ameghiniana 33 (4): 355–365. ISSN 1851-8044. ↑ Heredia, S., & Salgado, L. (2014). Posición estratigráfica de los estratos supracretácicos portadores de dinosaurios en Lago Pellegrini, Patagonia septentrional, Argentina. Ameghiniana, 36(2), 229-234.

musée Argentine Allen Anacleto +39
Figure 1. Evolution of macroecological traits in Dinosauria. Large scale event in dinosaur evolution (a); the origin of dinosaurs (star), hyperthermals (volcano), the earliest fossil Avialae (bird), the earliest fossil angiosperm (flower), the Cretaceous/Palaeogene mass extinction (asteroid). Phylogeny of dinosaurs (b) redrawn from Sereno and adapted to the current consensus and upon which an ancestral state reconstruction of temperature niche (mean annual temperature) after Chiarenza et al. is plotted; Mesozoic palaeogeographies (c) for Triassic (T), Jurassic (J) and Cretaceous (K). Silhouette colours symbolize body mass for each of the taxa represented; information on dietary habits are plotted after Barrett and Zanno & Makovicky; numbers represent clades discussed through this study: 1, Ornithischia; 2, Thyreophora; 3, Ornithopoda; 4, Hadrosauroidea; 5, Marginocephalia; 6, Ceratopsia; 7, Saurischia; 8, Sauropodomorpha; 9, Sauropoda; 10, Theropoda; 11, Ceratosauria; 12, Tetanurae; 13, Coelurosauria; 14, Maniraptoriformes; 15, Maniraptora; 16, Deinonychosauria; 17, Avialae; 18, Ornithothoraces. Palaeogeographies modified from original plots via R package ‘mapast’ using plate models by Scotese.

Figure 1. Evolution of macroecological traits in Dinosauria. Large scale event in dinosaur evolution (a); the origin of dinosaurs (star), hyperthermals (volcano), the earliest fossil Avialae (bird), the earliest fossil angiosperm (flower), the Cretaceous/Palaeogene mass extinction (asteroid). Phylogeny of dinosaurs (b) redrawn from Sereno and adapted to the current consensus and upon which an ancestral state reconstruction of temperature niche (mean annual temperature) after Chiarenza et al. is plotted; Mesozoic palaeogeographies (c) for Triassic (T), Jurassic (J) and Cretaceous (K). Silhouette colours symbolize body mass for each of the taxa represented; information on dietary habits are plotted after Barrett and Zanno & Makovicky; numbers represent clades discussed through this study: 1, Ornithischia; 2, Thyreophora; 3, Ornithopoda; 4, Hadrosauroidea; 5, Marginocephalia; 6, Ceratopsia; 7, Saurischia; 8, Sauropodomorpha; 9, Sauropoda; 10, Theropoda; 11, Ceratosauria; 12, Tetanurae; 13, Coelurosauria; 14, Maniraptoriformes; 15, Maniraptora; 16, Deinonychosauria; 17, Avialae; 18, Ornithothoraces. Palaeogeographies modified from original plots via R package ‘mapast’ using plate models by Scotese.

écaille Crétacé Jurassique Mésozoïque +23
Representative taxa from the Santonian Iharkút fauna from the Csehbánya Formation, Bakony Mountains, western Hungary. A Pannoniasaurus inexpectatus (Squamata, Mosasauroidea), dorsal vertebra (MTM uncatalogued) in dorsal view (photo by Réka Kalmár) B Foxemys trabanti (Pleurodira, Bothremydidae), skull (MTM V 2010.215.1.) in dorsal view (photo by Márton Rabi). C Bicuspidon aff. hatzegiensis (Squamata, Borioteiioidea), left dentary (MTM 2006.112.1.) in medial view (photo by László Makádi) D Basal tetanuran (Theropoda, Tetanurae), tooth (MTM V.01.54) in ?lingual view E Indeterminate abelisaurid (Theropoda, Abelisauridae), pedal ungual phalanx (MTM V 2008.43.1.) in lateral view F Pneumatoraptor fodori (Theropoda, Paraves), left scapulocoracoid (holotype, MTM V 2008.38.1.) in lateral view G Mochlodon vorosi (Ornithopoda, Rhabdodontidae), left dentary (holotype, MTM V 2010.105.1) in lateral view H Bakonydraco galaczi (Pterosauria, Azhdarchidae), mandible (holotype, MTM 2007.110.1) in dorsal view I Iharkutosuchus makadii (Eusuchia, Hylaeochampsidae), skull (holotype, MTM 2006.52.1) in dorsal view J Hungarosaurus tormai (Ankylosauria, Nodosauridae), right dentary (MTM 2007.25.2) in lateral view K Bauxitornis mindszentyae (Aves, Enantiornithes), left tarsometatarsus (holotype, MTM V 2009.38.1) in anterior view L Ajkaceratops kozmai (Ceratopsia), fused rostral and premaxillae (holotype, MTM V 2009.192.1) in lateral view. Scale bars: 2 cm in A, V, G, H, I, J; 1 cm in D, E, F, K, L; 1 mm in C.

Representative taxa from the Santonian Iharkút fauna from the Csehbánya Formation, Bakony Mountains, western Hungary. A Pannoniasaurus inexpectatus (Squamata, Mosasauroidea), dorsal vertebra (MTM uncatalogued) in dorsal view (photo by Réka Kalmár) B Foxemys trabanti (Pleurodira, Bothremydidae), skull (MTM V 2010.215.1.) in dorsal view (photo by Márton Rabi). C Bicuspidon aff. hatzegiensis (Squamata, Borioteiioidea), left dentary (MTM 2006.112.1.) in medial view (photo by László Makádi) D Basal tetanuran (Theropoda, Tetanurae), tooth (MTM V.01.54) in ?lingual view E Indeterminate abelisaurid (Theropoda, Abelisauridae), pedal ungual phalanx (MTM V 2008.43.1.) in lateral view F Pneumatoraptor fodori (Theropoda, Paraves), left scapulocoracoid (holotype, MTM V 2008.38.1.) in lateral view G Mochlodon vorosi (Ornithopoda, Rhabdodontidae), left dentary (holotype, MTM V 2010.105.1) in lateral view H Bakonydraco galaczi (Pterosauria, Azhdarchidae), mandible (holotype, MTM 2007.110.1) in dorsal view I Iharkutosuchus makadii (Eusuchia, Hylaeochampsidae), skull (holotype, MTM 2006.52.1) in dorsal view J Hungarosaurus tormai (Ankylosauria, Nodosauridae), right dentary (MTM 2007.25.2) in lateral view K Bauxitornis mindszentyae (Aves, Enantiornithes), left tarsometatarsus (holotype, MTM V 2009.38.1) in anterior view L Ajkaceratops kozmai (Ceratopsia), fused rostral and premaxillae (holotype, MTM V 2009.192.1) in lateral view. Scale bars: 2 cm in A, V, G, H, I, J; 1 cm in D, E, F, K, L; 1 mm in C.

dent vertèbre Hongrie Santonien +15
Seven ornithopods (top to bottom): Ouranosaurus, Tenontosaurus (background) with Convolosaurus (foreground), Muttaburrasaurus, Edmontosaurus annectens, Dryosaurus altus, Corythosaurus casuarius
Taxons Iguanodontia

Seven ornithopods (top to bottom): Ouranosaurus, Tenontosaurus (background) with Convolosaurus (foreground), Muttaburrasaurus, Edmontosaurus annectens, Dryosaurus altus, Corythosaurus casuarius

Clypeodonta Convolosaurus Corythosaurus Dryomorpha +9
Seven ornithopods (top to bottom): Ouranosaurus, Tenontosaurus (background) with Convolosaurus (foreground), Muttaburrasaurus, Edmontosaurus annectens, Dryosaurus altus, Corythosaurus casuarius
Taxons Euiguanodontia

Seven ornithopods (top to bottom): Ouranosaurus, Tenontosaurus (background) with Convolosaurus (foreground), Muttaburrasaurus, Edmontosaurus annectens, Dryosaurus altus, Corythosaurus casuarius

Clypeodonta Convolosaurus Corythosaurus Dryomorpha +9
Seven ornithopods (top to bottom): Ouranosaurus, Tenontosaurus (background) with Convolosaurus (foreground), Muttaburrasaurus, Edmontosaurus annectens, Dryosaurus altus, Corythosaurus casuarius
Taxons Dryomorpha

Seven ornithopods (top to bottom): Ouranosaurus, Tenontosaurus (background) with Convolosaurus (foreground), Muttaburrasaurus, Edmontosaurus annectens, Dryosaurus altus, Corythosaurus casuarius

Clypeodonta Convolosaurus Corythosaurus Dryomorpha +9
Seven ornithopods (top to bottom): Ouranosaurus, Tenontosaurus (background) with Convolosaurus (foreground), Muttaburrasaurus, Edmontosaurus annectens, Dryosaurus altus, Corythosaurus casuarius
Taxons Euornithopoda

Seven ornithopods (top to bottom): Ouranosaurus, Tenontosaurus (background) with Convolosaurus (foreground), Muttaburrasaurus, Edmontosaurus annectens, Dryosaurus altus, Corythosaurus casuarius

Clypeodonta Convolosaurus Corythosaurus Dryomorpha +9
Seven ornithopods (top to bottom): Ouranosaurus, Tenontosaurus (background) with Convolosaurus (foreground), Muttaburrasaurus, Edmontosaurus annectens, Dryosaurus altus, Corythosaurus casuarius
Taxons Clypeodonta

Seven ornithopods (top to bottom): Ouranosaurus, Tenontosaurus (background) with Convolosaurus (foreground), Muttaburrasaurus, Edmontosaurus annectens, Dryosaurus altus, Corythosaurus casuarius

Clypeodonta Convolosaurus Corythosaurus Dryomorpha +9
‘Koutalisaurus kohlerorum’ (Ornithopoda, Lambeosaurinae; indeterminate lambeosaurine sensu Prieto-Márquez et al. 2013), right dentary (IPS 29920 (formerly IPS SRA 27) in medial view (‘lower red unit’ of the Tremp Formation, Les Llaus near Sant Romà d’Abella, Lleida)

‘Koutalisaurus kohlerorum’ (Ornithopoda, Lambeosaurinae; indeterminate lambeosaurine sensu Prieto-Márquez et al. 2013), right dentary (IPS 29920 (formerly IPS SRA 27) in medial view (‘lower red unit’ of the Tremp Formation, Les Llaus near Sant Romà d’Abella, Lleida)

Koutalisaurus Lambeosaurinae Ornithopoda formation
Left femur of cf. Iguanodontidae gen. et sp. indet. (Ornithischia, Ornithopoda), IGP MZHLZ/2003/1, in posterior (A), lateral (B), medial (C), ante− rior (D), and distal (F) views, with transverse cross−section through the femoral shaft (E).

Left femur of cf. Iguanodontidae gen. et sp. indet. (Ornithischia, Ornithopoda), IGP MZHLZ/2003/1, in posterior (A), lateral (B), medial (C), ante− rior (D), and distal (F) views, with transverse cross−section through the femoral shaft (E).

Burianosaurus Iguanodontidae Ornithischia Ornithopoda