Kirtland

Skeletal reconstruction of Ahshislepelta minor, a small ankylosaur from the Late Cretaceous Kirtland Formation of New Mexico. While originally recovered as an ankylosaurid related to Gastonia, later analyses favored a nodosaurid position which this is based on. Based on the holotype SMP VP-1930, consisting of fragmentary shoulder, forelimb, and vertebral elements as well as several osteoderms. Unknown material filled in using Niobrarasaurus, Pawpawsaurus, Silvisaurus, Sauropelta, and Borealopelta. Total length is approximately 4.9 m through the centra. Notes: Osteoderm placement somewhat speculative, not all osteoderms and vertebral fragments are figured. References: Burns & Sullivan, 2011.

Sandstones of the lower Kirtland Formation (Late Campanian), outcropping near Coal Creek. New Mexico. Photo by Nick Longrich

Photograph of the holotype skull of Titanoceratops ouranos, from the late Campanian aged Kirtland/Fruitland beds of New Mexico, USA. Photograph by Nick Longrich

Comparison of cranial features between closely related southern Laramidian taxa; (A), Akainacephalus johnsoni (UMNH VP 20202) from the Late Cretaceous Kaiparowits Formation of Utah; and (B), Nodocephalosaurus kirtlandensis (SMP VP-900) from the Late Cretaceous Kirtland Formation of New Mexico, in left lateral views. Various synapomorphies are shared with N. kirtlandensis (highlighted in black and white arrows) and includes “flaring nostrils”; enlarged, laterally projecting, loreal osteoderms that are situated directly dorsal to the external nares. Other synapomorphies include pyramid-shaped nasal and frontal osteoderms positioned on the dorsal regions of the skull. A number of significant differences have been observed between both specimens; in A. johnsoni, the anterior, and posterior supraorbital bosses form an enlarged element that is somewhat backswept, whereas in N. kirtlandensis, the posterior and anterior supraorbital bosses are clearly defined as individual osteoderms, and are much smaller in size. Additionally, the squamosal horn in Akainacephalus is very small but is prominent and tetrahedrally shaped in Nodocephalosaurus. The quadratojugal horn in Akainacephalus is massive, has a subtriangular morphology in lateral view and projects almost entirely ventral, whereas in Nodocephalosaurus, the quadratojugal horn is smaller and has a typical fin-shaped morphology. Study sites: asob, anterior supraorbital boss; ext naris, external naris; laca, lacrimal caputegulum; loca, loreal caputegulum; naca, nasal caputegulae; orb, orbit; psob, posterior supraorbital boss; qjh, quadratojugal horn; sqh, squamosal horn.