humérus

Partie anatomique

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cast of Quetzalcoatlus northropi humerus 03 - Pterosaurs Flight in the Age of Dinosaurs

cast of Quetzalcoatlus northropi humerus 03 - Pterosaurs Flight in the Age of Dinosaurs

humérus vol États-Unis moulage +3
cast of Quetzalcoatlus northropi humerus 02 - Pterosaurs Flight in the Age of Dinosaurs

cast of Quetzalcoatlus northropi humerus 02 - Pterosaurs Flight in the Age of Dinosaurs

humérus vol États-Unis moulage +3
Cast of Quetzalcoatlus northropi humerus 01 - Pterosaurs Flight in the Age of Dinosaurs

Cast of Quetzalcoatlus northropi humerus 01 - Pterosaurs Flight in the Age of Dinosaurs

humérus vol moulage Dinosauria +2
Holotype specimen (PIMUZ A/III 1274) of Prosantosaurus scheffoldi gen. et spec. nov. from the upper Prosanto Formation (Early Ladinian, Middle Triassic) of Ducanfurgga locality no. 4, southwest of Davos, Canton of Grisons, south-eastern Switzerland. A Nearly complete specimen as prepared in dorsal view. The posterior part of the tail was lost prior to burial. Both forearms are not visible but lie below the trunk region, pointing in an anteromedial direction (see Additional file 1: Fig. S3A). B Detail of skull and anterior neck region. C Outline sketch of skull sutures. D Detail of shoulder girdle (claviculae, scapulae) and anterior dorsal vertebrae and ribs. E Detail of right humerus. F Detail of posterior dorsal vertebrae and ribs, sacral vertebrae and ribs, and anterior caudal vertebrae and ribs. G Detail of left ilium and hindlimb. ar articular; as astragalus; bo basioccipital; cal calcaneus; cl clavicula; co coracoid; d dentary; en external naris; eo exoccipital; fe femur; fi fibula; fr frontal; hu humerus; il ilium; in internal naris; is ischium; j jugal; mx maxilla; na naris; o orbit; pa parietal; pl palatine; pmx premaxilla; pof postfrontal; po postorbital; prf prefrontal; pt pterygoid; pu pubis; q quadrate; qj quadratojugal; ti tibia; sacr sacral rib; sc scapula; so supraoccipital; sp splenial; sq squamosal; su surangular; utf upper temporal fenestra; v vomer
Taxons Prosantosaurus

Holotype specimen (PIMUZ A/III 1274) of Prosantosaurus scheffoldi gen. et spec. nov. from the upper Prosanto Formation (Early Ladinian, Middle Triassic) of Ducanfurgga locality no. 4, southwest of Davos, Canton of Grisons, south-eastern Switzerland. A Nearly complete specimen as prepared in dorsal view. The posterior part of the tail was lost prior to burial. Both forearms are not visible but lie below the trunk region, pointing in an anteromedial direction (see Additional file 1: Fig. S3A). B Detail of skull and anterior neck region. C Outline sketch of skull sutures. D Detail of shoulder girdle (claviculae, scapulae) and anterior dorsal vertebrae and ribs. E Detail of right humerus. F Detail of posterior dorsal vertebrae and ribs, sacral vertebrae and ribs, and anterior caudal vertebrae and ribs. G Detail of left ilium and hindlimb. ar articular; as astragalus; bo basioccipital; cal calcaneus; cl clavicula; co coracoid; d dentary; en external naris; eo exoccipital; fe femur; fi fibula; fr frontal; hu humerus; il ilium; in internal naris; is ischium; j jugal; mx maxilla; na naris; o orbit; pa parietal; pl palatine; pmx premaxilla; pof postfrontal; po postorbital; prf prefrontal; pt pterygoid; pu pubis; q quadrate; qj quadratojugal; ti tibia; sacr sacral rib; sc scapula; so supraoccipital; sp splenial; sq squamosal; su surangular; utf upper temporal fenestra; v vomer

humérus Suisse Ladinien Trias moyen +6
Diagnostic skeletal elements of the type specimen of Franconiasaurus brevispinus gen. et sp. nov. (BT 011224.00). (A) main slab showing partly articulated skeleton in dorsal view. Posterior part of right mandibular ramus in (B) lateral and (C) dorsal aspect; anterior cervical vertebra (D) in articular and (E) lateral view; (F) posterior cervical vertebra in lateral view; (G) cervical rib in dorsolateral view; (H) dorsal vertebra in posterior view; (I) left scapula in medial view; (J) right coracoid in supposed dorsal view; (K) left humerus in dorsal view; (L) left femur with adjacent epipodials in dorsal view; and (M) left ilium in lateral view.
Taxons Franconiasaurus

Diagnostic skeletal elements of the type specimen of Franconiasaurus brevispinus gen. et sp. nov. (BT 011224.00). (A) main slab showing partly articulated skeleton in dorsal view. Posterior part of right mandibular ramus in (B) lateral and (C) dorsal aspect; anterior cervical vertebra (D) in articular and (E) lateral view; (F) posterior cervical vertebra in lateral view; (G) cervical rib in dorsolateral view; (H) dorsal vertebra in posterior view; (I) left scapula in medial view; (J) right coracoid in supposed dorsal view; (K) left humerus in dorsal view; (L) left femur with adjacent epipodials in dorsal view; and (M) left ilium in lateral view.

humérus vertèbre spécimen Franconiasaurus +1
MUHNCAL.20173, holotype specimen of Gondwananectes osvaldoi. A, quarry scheme of the recovered blocks (numbered 1–7). Blocks 1–3 preserve contact surfaces and they are arranged consistently. The position of blocks 4–7 is tentative due to the lack of contact surfaces, although it is based on the anatomical elements preserved. B, schematic diagram of the preserved anatomical elements in MUHNCAL.20173. Abbreviations: c1–c10, cervical centra/vertebrae; d1–d3, dorsal centra; d1na–d3na, neural arches of the d1–d3 dorsal centra; il, ilium fragment; in, indeterminate; lco, left coracoid, lcna, last cervical neural arch; lh, left humerus; lsc, left scapula (ventral process); p1–p2, pectoral centra; pd, posterior dorsal centrum; pna, pectoral neural arch; r, ribs; rh, right humerus (distal); rsc, right scapula (dorsal process). Scale bars represent 10 cm.
Taxons Gondwananectes

MUHNCAL.20173, holotype specimen of Gondwananectes osvaldoi. A, quarry scheme of the recovered blocks (numbered 1–7). Blocks 1–3 preserve contact surfaces and they are arranged consistently. The position of blocks 4–7 is tentative due to the lack of contact surfaces, although it is based on the anatomical elements preserved. B, schematic diagram of the preserved anatomical elements in MUHNCAL.20173. Abbreviations: c1–c10, cervical centra/vertebrae; d1–d3, dorsal centra; d1na–d3na, neural arches of the d1–d3 dorsal centra; il, ilium fragment; in, indeterminate; lco, left coracoid, lcna, last cervical neural arch; lh, left humerus; lsc, left scapula (ventral process); p1–p2, pectoral centra; pd, posterior dorsal centrum; pna, pectoral neural arch; r, ribs; rh, right humerus (distal); rsc, right scapula (dorsal process). Scale bars represent 10 cm.

humérus écaille holotype spécimen +1
New wing skeleton of Forfexopterus (SDUST-V1003) from Jiufotang Formation of Early Cretaceous Jehol Biota in Jianchang, western Liaoning, northeastern China. Abbreviations: ca, carpus; co, coracoid portion; hu, humerus; mc, metacarpals I–IV; mdI–III, manual digits I–III; pt, pteroid; ra, radius; sc, scapular portion; ul, ulna; wp1–4, wing phalanges 1–4.
Taxons Forfexopterus

New wing skeleton of Forfexopterus (SDUST-V1003) from Jiufotang Formation of Early Cretaceous Jehol Biota in Jianchang, western Liaoning, northeastern China. Abbreviations: ca, carpus; co, coracoid portion; hu, humerus; mc, metacarpals I–IV; mdI–III, manual digits I–III; pt, pteroid; ra, radius; sc, scapular portion; ul, ulna; wp1–4, wing phalanges 1–4.

humérus Chine Jiufotang Crétacé +4
Photograph (A) and line drawings (B) of Gladocephaloideus jingangshanensis (JPM 2014–004). Abbreviations: ca, carpals; cr, coracoids; cv, cervical vertebrae; d, dentray; dg, deep groove along the mid-line of the mandibular symphysis; dv, dorsal vertebrae; dr, dorsal ribs; etp, extensor tendon process; f, frontal; fc, fifth carpal; fe, femur; fi, fibula; fin, fingers; h, humerus; il, ilium; m, maxilla; mmttsI-IV, metatrals I-IV; mttv, metatarsal V; nao, nasoantorbital opening; or, orbital; pcr, parietal crest; pm, premaxilla; pt, pteroid; ra, radius; rdl, radiale; sc, scapula; st, sternum; sl, sclerotic rings; t, teeth; tc, tooth sockets; ti, tibia; ul, ulna; wm, wing metacarpal; wph1-4, wing phalanges 1–4. Scale bar = 5 cm.
Taxons Gladocephaloideus

Photograph (A) and line drawings (B) of Gladocephaloideus jingangshanensis (JPM 2014–004). Abbreviations: ca, carpals; cr, coracoids; cv, cervical vertebrae; d, dentray; dg, deep groove along the mid-line of the mandibular symphysis; dv, dorsal vertebrae; dr, dorsal ribs; etp, extensor tendon process; f, frontal; fc, fifth carpal; fe, femur; fi, fibula; fin, fingers; h, humerus; il, ilium; m, maxilla; mmttsI-IV, metatrals I-IV; mttv, metatarsal V; nao, nasoantorbital opening; or, orbital; pcr, parietal crest; pm, premaxilla; pt, pteroid; ra, radius; rdl, radiale; sc, scapula; st, sternum; sl, sclerotic rings; t, teeth; tc, tooth sockets; ti, tibia; ul, ulna; wm, wing metacarpal; wph1-4, wing phalanges 1–4. Scale bar = 5 cm.

crête humérus écaille dent +2
FSAC-OB 7, holotype right humerus, Simurghia robustus.
In (A), dorsal view, (B) ventral view, and (C) posterior view. Abbreviations: dpc, deltopectoral crest; msc, muscle scar; scpr, supracondylar process of the ectepicondyle; vp, ventral pillar; vt, ventral tubercle of the deltopectoral crest.
Taxons Simurghia

FSAC-OB 7, holotype right humerus, Simurghia robustus. In (A), dorsal view, (B) ventral view, and (C) posterior view. Abbreviations: dpc, deltopectoral crest; msc, muscle scar; scpr, supracondylar process of the ectepicondyle; vp, ventral pillar; vt, ventral tubercle of the deltopectoral crest.

crête humérus holotype Simurghia
FSAC-OB 232, holotype skeleton of Barbaridactylus grandis.
(A) Left humerus in anterior view, (B) right femur in anterior view, (C) right radius and ulna in posterior view, (D) cervical vertebra in ventral view, (E) left scapulocoracoid in medial view, and (F) posterior ramus of the right mandible in medial view.

Abbreviations: co, coracoid; cot, cotyle of mandible; cvr, cervical ribs; dm, dorsal margin of mandible; dpc, deltopectoral crest; gl, glenoid; fh, femoral head; gt, greater trochanter; hh, humeral head; hyp, hypapophysis; olp, olecranon process; pex, postexapophysis; pn, pneumatopore; rad, radius; sca, scapula; scp, supracondylar process of the ectepicondyle; uc, ulnar crest; uln, ulna; vm, ventral margin of mandible.
Taxons Barbaridactylus

FSAC-OB 232, holotype skeleton of Barbaridactylus grandis. (A) Left humerus in anterior view, (B) right femur in anterior view, (C) right radius and ulna in posterior view, (D) cervical vertebra in ventral view, (E) left scapulocoracoid in medial view, and (F) posterior ramus of the right mandible in medial view. Abbreviations: co, coracoid; cot, cotyle of mandible; cvr, cervical ribs; dm, dorsal margin of mandible; dpc, deltopectoral crest; gl, glenoid; fh, femoral head; gt, greater trochanter; hh, humeral head; hyp, hypapophysis; olp, olecranon process; pex, postexapophysis; pn, pneumatopore; rad, radius; sca, scapula; scp, supracondylar process of the ectepicondyle; uc, ulnar crest; uln, ulna; vm, ventral margin of mandible.

crête humérus vertèbre holotype +2
Mimodactylus libanensis gen. et sp. nov. (a) Photo and drawing of the complete specimen. (b) Close up of scapula and coracoid. (c) Detail of the wrist, showing the relation of the pteroid and the carpus. (d) Detail of the humerus. Scale-bars, a: 50 mm; b-d: 10 mm. Abbreviations. car: carpus; cdv: caudal vertebrae; cor: coracoid; cra: cranium; cs: cristospine; cv: cervical vertebrae; d: dentary; dcar: distal carpals; dpc: deltopectoral crest; dri: dorsal ribs; dv: dorsal vertebrae; fe: femur; hu: humerus; man: mandible; mcI-III: first to third metacarpals; mcIV: wing metacarpal; mtar: metatarsals; pcar: proximal carpals; ph1d4: first wing phalanx; ph2d4: second wing phalanx; ph3d4: third wing phalanx; ph4d4: fourth wing phalanx; ppu: prepubis; ptd: pteroid; ra: radius; sca: scapula; sri: sacral ribs; stp: sternal plate; sym: mandibular symphysis; tar: tarsus; te: teeth; ti: tibia; ul: ulna; the abbreviations ‘l’ and ‘r’ represents respectively left and right.
Taxons Mimodactylus

Mimodactylus libanensis gen. et sp. nov. (a) Photo and drawing of the complete specimen. (b) Close up of scapula and coracoid. (c) Detail of the wrist, showing the relation of the pteroid and the carpus. (d) Detail of the humerus. Scale-bars, a: 50 mm; b-d: 10 mm. Abbreviations. car: carpus; cdv: caudal vertebrae; cor: coracoid; cra: cranium; cs: cristospine; cv: cervical vertebrae; d: dentary; dcar: distal carpals; dpc: deltopectoral crest; dri: dorsal ribs; dv: dorsal vertebrae; fe: femur; hu: humerus; man: mandible; mcI-III: first to third metacarpals; mcIV: wing metacarpal; mtar: metatarsals; pcar: proximal carpals; ph1d4: first wing phalanx; ph2d4: second wing phalanx; ph3d4: third wing phalanx; ph4d4: fourth wing phalanx; ppu: prepubis; ptd: pteroid; ra: radius; sca: scapula; sri: sacral ribs; stp: sternal plate; sym: mandibular symphysis; tar: tarsus; te: teeth; ti: tibia; ul: ulna; the abbreviations ‘l’ and ‘r’ represents respectively left and right.

crête humérus écaille dessin +3
Aralazhdarcho bostobensis, ZIN PH 57/43, a proximal fragment of a left humerus in proximal (a), ventral (b), anterior (c), dorsal (d), and posterior (e) views. This specimen is from the Shakh Shakh II locality in the northeasten Aral Sea region of Kazakhstan; Bostobe Formation, Upper Cretaceous (Santonian – lower Campanian). Abbreviations: h, humeral head; pf, pneumatic foramen; uc, ulnar crest. Scale bar is 10 mm.
Taxons Aralazhdarcho

Aralazhdarcho bostobensis, ZIN PH 57/43, a proximal fragment of a left humerus in proximal (a), ventral (b), anterior (c), dorsal (d), and posterior (e) views. This specimen is from the Shakh Shakh II locality in the northeasten Aral Sea region of Kazakhstan; Bostobe Formation, Upper Cretaceous (Santonian – lower Campanian). Abbreviations: h, humeral head; pf, pneumatic foramen; uc, ulnar crest. Scale bar is 10 mm.

crête humérus écaille Kazakhstan +7
Entire skeleton of Jidapterus edentus (RCPS-030366CY) preserved in a slab mostly in right lateral view.

Abbreviations: aa, the atlas-axis complex; bo, basioccipital; br, braincase; cve, cervical vertebrae; cv3, the third cervical vertebra; cv8, the eighth cervical vertebra; ds, dentaries; dv, dorsal vertebra; dve, dorsal vertebrae; ep, epiphysis; j, jugal; lc, left carpus; lf, left femur; li, left ilium; lis, left ischium; lm, left manual digits; lp, left pubis; lpe, left pes; lpt, left pteroid; lr, left radius; lsc, left scapula-coracoid; lt, left tibia; lta, left tarsus; lu, left ulna; lwmc, left wing metacarpal; lwp1–4; left wing phalanges 1–4; p, pterygoid; pp, postacetabular process of the right ilium; q, quadrate; rc, right carpus; rh, right humerus; rf, right femur; rfi, right fibula; ribs, ribs; ris, right ischium; rm, right manual digits; rmcI, right metacarpal I; rp, right pubis; rpe, right pes; rpt, right pteroid; rr, right radius; rsc, right scapula-coracoid; rt, right tibia; ru, right ulna; rwmc, right wing metacarpal; rwp1–4, right wing phalanges 1–4; rostrum, rostrum; st, sternum; sv, sacral vertebra; sve, sacral vertebrae. Gray shaded areas bound by solid lines indicate preserved bones, dashed lines indicate bones with margins discerned by impression, and lack of lines indicate missing bone or displacement along cracks in the slab.
Taxons Jidapterus

Entire skeleton of Jidapterus edentus (RCPS-030366CY) preserved in a slab mostly in right lateral view. Abbreviations: aa, the atlas-axis complex; bo, basioccipital; br, braincase; cve, cervical vertebrae; cv3, the third cervical vertebra; cv8, the eighth cervical vertebra; ds, dentaries; dv, dorsal vertebra; dve, dorsal vertebrae; ep, epiphysis; j, jugal; lc, left carpus; lf, left femur; li, left ilium; lis, left ischium; lm, left manual digits; lp, left pubis; lpe, left pes; lpt, left pteroid; lr, left radius; lsc, left scapula-coracoid; lt, left tibia; lta, left tarsus; lu, left ulna; lwmc, left wing metacarpal; lwp1–4; left wing phalanges 1–4; p, pterygoid; pp, postacetabular process of the right ilium; q, quadrate; rc, right carpus; rh, right humerus; rf, right femur; rfi, right fibula; ribs, ribs; ris, right ischium; rm, right manual digits; rmcI, right metacarpal I; rp, right pubis; rpe, right pes; rpt, right pteroid; rr, right radius; rsc, right scapula-coracoid; rt, right tibia; ru, right ulna; rwmc, right wing metacarpal; rwp1–4, right wing phalanges 1–4; rostrum, rostrum; st, sternum; sv, sacral vertebra; sve, sacral vertebrae. Gray shaded areas bound by solid lines indicate preserved bones, dashed lines indicate bones with margins discerned by impression, and lack of lines indicate missing bone or displacement along cracks in the slab.

humérus vertèbre Chaoyangopteridae Chaoyangopterinae +2
Entire skeleton of Jidapterus edentus (RCPS-030366CY) preserved in a slab mostly in right lateral view.

Abbreviations: aa, the atlas-axis complex; bo, basioccipital; br, braincase; cve, cervical vertebrae; cv3, the third cervical vertebra; cv8, the eighth cervical vertebra; ds, dentaries; dv, dorsal vertebra; dve, dorsal vertebrae; ep, epiphysis; j, jugal; lc, left carpus; lf, left femur; li, left ilium; lis, left ischium; lm, left manual digits; lp, left pubis; lpe, left pes; lpt, left pteroid; lr, left radius; lsc, left scapula-coracoid; lt, left tibia; lta, left tarsus; lu, left ulna; lwmc, left wing metacarpal; lwp1–4; left wing phalanges 1–4; p, pterygoid; pp, postacetabular process of the right ilium; q, quadrate; rc, right carpus; rh, right humerus; rf, right femur; rfi, right fibula; ribs, ribs; ris, right ischium; rm, right manual digits; rmcI, right metacarpal I; rp, right pubis; rpe, right pes; rpt, right pteroid; rr, right radius; rsc, right scapula-coracoid; rt, right tibia; ru, right ulna; rwmc, right wing metacarpal; rwp1–4, right wing phalanges 1–4; rostrum, rostrum; st, sternum; sv, sacral vertebra; sve, sacral vertebrae. Gray shaded areas bound by solid lines indicate preserved bones, dashed lines indicate bones with margins discerned by impression, and lack of lines indicate missing bone or displacement along cracks in the slab.
Taxons Chaoyangopterinae

Entire skeleton of Jidapterus edentus (RCPS-030366CY) preserved in a slab mostly in right lateral view. Abbreviations: aa, the atlas-axis complex; bo, basioccipital; br, braincase; cve, cervical vertebrae; cv3, the third cervical vertebra; cv8, the eighth cervical vertebra; ds, dentaries; dv, dorsal vertebra; dve, dorsal vertebrae; ep, epiphysis; j, jugal; lc, left carpus; lf, left femur; li, left ilium; lis, left ischium; lm, left manual digits; lp, left pubis; lpe, left pes; lpt, left pteroid; lr, left radius; lsc, left scapula-coracoid; lt, left tibia; lta, left tarsus; lu, left ulna; lwmc, left wing metacarpal; lwp1–4; left wing phalanges 1–4; p, pterygoid; pp, postacetabular process of the right ilium; q, quadrate; rc, right carpus; rh, right humerus; rf, right femur; rfi, right fibula; ribs, ribs; ris, right ischium; rm, right manual digits; rmcI, right metacarpal I; rp, right pubis; rpe, right pes; rpt, right pteroid; rr, right radius; rsc, right scapula-coracoid; rt, right tibia; ru, right ulna; rwmc, right wing metacarpal; rwp1–4, right wing phalanges 1–4; rostrum, rostrum; st, sternum; sv, sacral vertebra; sve, sacral vertebrae. Gray shaded areas bound by solid lines indicate preserved bones, dashed lines indicate bones with margins discerned by impression, and lack of lines indicate missing bone or displacement along cracks in the slab.

humérus vertèbre Chaoyangopteridae Chaoyangopterinae +2
Nemicolopterus crypticus holotype (IVPP V-14377).

(A) Skeleton overview, and (B) schematic drawing. (C) Skull (right lateral view), and (D) schematic drawing. Abbreviations: cdv, caudal vertebrae; co, coracoid; cv, cervical vertebra; d1–d4, digits 1–4; f, frontal; fe, femur; h, humerus; hy, hyoid; il, illium; is, ischium; j, jugal; l, left; la, lacrimal; mand, mandible; mc, metacarpal; mt, metatarsal; naof, nasoantorbital fenestra; or, orbit; pa, parietal; pm, premaxilla; ph, phalanx; ti, tibia; r, right; sca, scapula. Scale bars: A–B, 100 mm; C–D, 5 mm.
Taxons Nemicolopterus

Nemicolopterus crypticus holotype (IVPP V-14377). (A) Skeleton overview, and (B) schematic drawing. (C) Skull (right lateral view), and (D) schematic drawing. Abbreviations: cdv, caudal vertebrae; co, coracoid; cv, cervical vertebra; d1–d4, digits 1–4; f, frontal; fe, femur; h, humerus; hy, hyoid; il, illium; is, ischium; j, jugal; l, left; la, lacrimal; mand, mandible; mc, metacarpal; mt, metatarsal; naof, nasoantorbital fenestra; or, orbit; pa, parietal; pm, premaxilla; ph, phalanx; ti, tibia; r, right; sca, scapula. Scale bars: A–B, 100 mm; C–D, 5 mm.

humérus écaille vertèbre dessin +4
Selected elements used in the diagnosis of Saltriovenator zanellai n. gen. n. sp.

Right humerus in medial (A), frontal (B) and distal (C) views; (D) left scapula, medial view; (E) right scapular glenoid and coracoid, lateral view; (F) furcula, ventral view; tooth, labial (G) and apical (H) views; (I) left humerus, medial view; right second metacarpal in dorsal (J), lateral (L) and distal (N) views; first phalanx of the right second digit in dorsal (K), lateral (M) and proximal (O) views; (P–T) right third digit in proximal, dorsal and lateral views; (U) right distal tarsal IV, proximal view; third right metatarsal in proximal (V) and frontal (X) views; second right metatarsal, proximal (W) and frontal (Y) views; (Z) reconstructed skeleton showing identified elements (red). Abbreviations as in text, asterisks mark autapomorphic traits. Scale bars: 10 cm in (A)–(E), (I), and (U)–(Y); two cm in (F), and (J)–(T); one cm in (G).
Taxons Saltriovenator

Selected elements used in the diagnosis of Saltriovenator zanellai n. gen. n. sp. Right humerus in medial (A), frontal (B) and distal (C) views; (D) left scapula, medial view; (E) right scapular glenoid and coracoid, lateral view; (F) furcula, ventral view; tooth, labial (G) and apical (H) views; (I) left humerus, medial view; right second metacarpal in dorsal (J), lateral (L) and distal (N) views; first phalanx of the right second digit in dorsal (K), lateral (M) and proximal (O) views; (P–T) right third digit in proximal, dorsal and lateral views; (U) right distal tarsal IV, proximal view; third right metatarsal in proximal (V) and frontal (X) views; second right metatarsal, proximal (W) and frontal (Y) views; (Z) reconstructed skeleton showing identified elements (red). Abbreviations as in text, asterisks mark autapomorphic traits. Scale bars: 10 cm in (A)–(E), (I), and (U)–(Y); two cm in (F), and (J)–(T); one cm in (G).

humérus écaille dent dessin +2
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