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Skeletal reconstructions of Dinosaur Park Formation caenagnathids. Skeletal reconstructions of Citipes elegans (left), Chirostenotes pergracilis (middle), and Caenagnathus collinsi (right), showing variation in skeletal representation and body size. Previously referred material is indicated in white and newly referred material is indicated in red for each taxon. Blue asterisks indicate elements that have been histologically sampled for each taxon. Citipes elegans: dentaries, metatarsal IV; Chirostenotes pergracilis: dentaries, tibia; Caenagnathus collinsi: pubis.

Skeletal reconstructions of Dinosaur Park Formation caenagnathids. Skeletal reconstructions of Citipes elegans (left), Chirostenotes pergracilis (middle), and Caenagnathus collinsi (right), showing variation in skeletal representation and body size. Previously referred material is indicated in white and newly referred material is indicated in red for each taxon. Blue asterisks indicate elements that have been histologically sampled for each taxon. Citipes elegans: dentaries, metatarsal IV; Chirostenotes pergracilis: dentaries, tibia; Caenagnathus collinsi: pubis.

Dinosaur Park Caenagnathidae Caenagnathus Chirostenotes +4
Tarsometatarsi of caenagnathids from the Upper Campanian Dinosaur Park Formation, Dinosaur Provincial Park, Alberta, Canada. A. TMP 1993.036.0181 (Chirostenotes pergracilis). B. TMP 1982.016.0006 (Citipes elegans). C. ROM 781 (Citipes elegans holotype). D. TMP 1996.012.0141 (Citipes elegans). Tarsometatarsi in proximal (A1–D1), anterior (A2–D2), and posterior (A3–D3) views.

Tarsometatarsi of caenagnathids from the Upper Campanian Dinosaur Park Formation, Dinosaur Provincial Park, Alberta, Canada. A. TMP 1993.036.0181 (Chirostenotes pergracilis). B. TMP 1982.016.0006 (Citipes elegans). C. ROM 781 (Citipes elegans holotype). D. TMP 1996.012.0141 (Citipes elegans). Tarsometatarsi in proximal (A1–D1), anterior (A2–D2), and posterior (A3–D3) views.

Canada Dinosaur Park Campanien holotype +6
Original figure caption: Trackway S1 (Eosauropus sp.), here attributed to a sauropod trackmaker based on pedal synapomorphies; trackmaker is moving towards the south-west. Two consequtive pes impressions of a tridactyl Grallator [i.e. a theropod] trackway can be seen left to the S1 trackway.
Note: The tracks are preserved on a bedding plane of a thin siltstone bed of the Late Triassic Fleming Fjord Formation of East Greenland. A) shows a photograph of the trackway(s) as preserved on the bedding plane (i.e. as concave epireliefs); B) shows a color shaded relief map based on a high-resolution photogrammetric 3D-model of the bedding plane; C) is an interpretative outline drawing of the S1 trackway; abbreviations: LM = left manus (i.e. forefoot), LP = left pes (i.e. hindfoot), RM = right manus, RP = right pes, numbers increase in walking direction.

Original figure caption: Trackway S1 (Eosauropus sp.), here attributed to a sauropod trackmaker based on pedal synapomorphies; trackmaker is moving towards the south-west. Two consequtive pes impressions of a tridactyl Grallator [i.e. a theropod] trackway can be seen left to the S1 trackway. Note: The tracks are preserved on a bedding plane of a thin siltstone bed of the Late Triassic Fleming Fjord Formation of East Greenland. A) shows a photograph of the trackway(s) as preserved on the bedding plane (i.e. as concave epireliefs); B) shows a color shaded relief map based on a high-resolution photogrammetric 3D-model of the bedding plane; C) is an interpretative outline drawing of the S1 trackway; abbreviations: LM = left manus (i.e. forefoot), LP = left pes (i.e. hindfoot), RM = right manus, RP = right pes, numbers increase in walking direction.

dessin Groenland Trias supérieur Trias +4
Drawings of Dryolestida left upper−molar, modified from the original descriptions. A–C, Kimmeridgian; D–F, Tithonian–Berriasian; G–H, early Barremian; I, late Barremian. A. Left M4 or M5 of Comotherium richi Prothero, 1981, from the Upper Jurassic Morrison Formation, Como Bluff locality, Wyoming, USA. B. Right M5, reversed, of Dryolestes leiriensis Martin, 1999, from Guimarota, Portugal.C. Left M6 of Krebsotherium lusitanicum Martin, 1999, from Guimarota, Portugal. D. Right M4 or M 6 (reversed) of Portopinheirodon asymmetricus Martin, 1999, Porto Pinheiro, Lourinha, Portugal. E. Right M5 (reversed) of Laolestes andresi Martin, 1999, from Porto Pinheiro, Lourinha, Portugal. F. Right upper molar of Donodon perscriptoris Sigogneau−Russell, 1991, Anoual, Morocco.G. Left M4 orM5 of Crusafontia amoae sp. nov., holotype, Cuesta Corrales 2, El Castellar Formation, Galve, Teruel, Spain. H. Right M6 orM7 (reversed) of Crusafontia amoae sp. nov., first described as upper molar of Crusafontia cuencana by Krebs (1993), P−2 H4 Pelejón 2, Galve, Teruel, Spain. I. Left M2 or M3, of Crusafontia cuencana Henkel and Krebs, 1969, Uña, La Huérguina Formation, Cuenca, Spain, (from Krebs 1993). Scale bars 1 mm.

Drawings of Dryolestida left upper−molar, modified from the original descriptions. A–C, Kimmeridgian; D–F, Tithonian–Berriasian; G–H, early Barremian; I, late Barremian. A. Left M4 or M5 of Comotherium richi Prothero, 1981, from the Upper Jurassic Morrison Formation, Como Bluff locality, Wyoming, USA. B. Right M5, reversed, of Dryolestes leiriensis Martin, 1999, from Guimarota, Portugal.C. Left M6 of Krebsotherium lusitanicum Martin, 1999, from Guimarota, Portugal. D. Right M4 or M 6 (reversed) of Portopinheirodon asymmetricus Martin, 1999, Porto Pinheiro, Lourinha, Portugal. E. Right M5 (reversed) of Laolestes andresi Martin, 1999, from Porto Pinheiro, Lourinha, Portugal. F. Right upper molar of Donodon perscriptoris Sigogneau−Russell, 1991, Anoual, Morocco.G. Left M4 orM5 of Crusafontia amoae sp. nov., holotype, Cuesta Corrales 2, El Castellar Formation, Galve, Teruel, Spain. H. Right M6 orM7 (reversed) of Crusafontia amoae sp. nov., first described as upper molar of Crusafontia cuencana by Krebs (1993), P−2 H4 Pelejón 2, Galve, Teruel, Spain. I. Left M2 or M3, of Crusafontia cuencana Henkel and Krebs, 1969, Uña, La Huérguina Formation, Cuenca, Spain, (from Krebs 1993). Scale bars 1 mm.

description dessin Maroc Portugal +11
Holotype specimen TMP 2000.29.01 of the ophthalmosaurian ichthyosaur Athabascasaurus bitumineus from the Lower Cretaceous Clearwater Formation of Alberta, in Royal Tyrrell Museum, Drumheller, Alberta, Canada.
Taxons Athabascasaurus

Holotype specimen TMP 2000.29.01 of the ophthalmosaurian ichthyosaur Athabascasaurus bitumineus from the Lower Cretaceous Clearwater Formation of Alberta, in Royal Tyrrell Museum, Drumheller, Alberta, Canada.

musée Canada Crétacé holotype +5
Fig 2. Chrono-, bio- and lithostratigraphic context of the Reuchenette Formation in the Ajoie district, Canton Jura, NW Switzerland (modified from [20,23,28,31]. Four track-bearing intervals, named lower, intermediate, and upper (dinosaur track) levels, and track levels 600 have been identified within the Courtedoux Member (Nerinean Limestones, sensu [32]). All studied material comes from the intermediate (levels 1000–1100) and upper (levels 1500–1650) dinosaur track levels, details shown on inset on the upper right.

Fig 2. Chrono-, bio- and lithostratigraphic context of the Reuchenette Formation in the Ajoie district, Canton Jura, NW Switzerland (modified from [20,23,28,31]. Four track-bearing intervals, named lower, intermediate, and upper (dinosaur track) levels, and track levels 600 have been identified within the Courtedoux Member (Nerinean Limestones, sensu [32]). All studied material comes from the intermediate (levels 1000–1100) and upper (levels 1500–1650) dinosaur track levels, details shown on inset on the upper right.

Suisse Reuchenette Dinosauria Megalosauripus +1
Remake of the original picture of the Drzewica Formation. Terrestrial environment of the Pliensbachian-Toarcian boundary of Fennoscandinavia  Inland environment of the  Bornholm Coast, nearby the German realm of the Ciechocinek Formation. Includes
Ciechocinek Formation (Lower Toarcian, Bones) and Drzwica Formation (Latest Pliensbachian, Footprints) Fauna
Sorthat Formation environment, fluvial influenced mainland with Cheirolepidaceae and Bennetitales as dominant flora
Dinosaurs are based on material found on various locations of Northern Germany, and Footprints of the underliying Drzewica Formation at the Holy Cross Mountains, connected with Bornholm at the time.
Dinosaur Species appeared: 

Megalosauripus isp. Large Footprints (+65 cm) found on the Drzewica Formation. There is a dorsal vertebrae on the German Margin of the Ciechocinek Formation assigend to Megalosauria (Huene, 1966).
Gravisauria spp. representing the Grimmen Sauropod reported on 2014, as a taxon related with Tazoudasaurus. Barapasaurus-like footprints are know from the Drzewica Formation.
Coelophysoidea spp. based on coeval Anchisauripus tracks from the Holy Cross Mountains.
Basal Ornithischan, related to Eocursor, based on a crouching trace (Gerard Dariusz Gierlinski, Martin G. Lockley, Grzegorz Niedźwiedzki:2009).
Massospondylidae spp. based on Otozum-like tracks.

Remake of the original picture of the Drzewica Formation. Terrestrial environment of the Pliensbachian-Toarcian boundary of Fennoscandinavia Inland environment of the Bornholm Coast, nearby the German realm of the Ciechocinek Formation. Includes Ciechocinek Formation (Lower Toarcian, Bones) and Drzwica Formation (Latest Pliensbachian, Footprints) Fauna Sorthat Formation environment, fluvial influenced mainland with Cheirolepidaceae and Bennetitales as dominant flora Dinosaurs are based on material found on various locations of Northern Germany, and Footprints of the underliying Drzewica Formation at the Holy Cross Mountains, connected with Bornholm at the time. Dinosaur Species appeared: Megalosauripus isp. Large Footprints (+65 cm) found on the Drzewica Formation. There is a dorsal vertebrae on the German Margin of the Ciechocinek Formation assigend to Megalosauria (Huene, 1966). Gravisauria spp. representing the Grimmen Sauropod reported on 2014, as a taxon related with Tazoudasaurus. Barapasaurus-like footprints are know from the Drzewica Formation. Coelophysoidea spp. based on coeval Anchisauripus tracks from the Holy Cross Mountains. Basal Ornithischan, related to Eocursor, based on a crouching trace (Gerard Dariusz Gierlinski, Martin G. Lockley, Grzegorz Niedźwiedzki:2009). Massospondylidae spp. based on Otozum-like tracks.

os Allemagne Pliensbachien Toarcien +7
Dentary of Echinodon becklesii from the Lower Cretaceous Purbeck Formation of England. Left dentary (NHMUK 48215b) in lateral (A), medial (B), and dorsal (C) views.
Taxons Echinodon

Dentary of Echinodon becklesii from the Lower Cretaceous Purbeck Formation of England. Left dentary (NHMUK 48215b) in lateral (A), medial (B), and dorsal (C) views.

Crétacé Echinodon Sauraechmodon formation
Dentary of Echinodon becklesii from the Lower Cretaceous Purbeck Formation of England. Left dentary (NHMUK 48215b) in lateral (A), medial (B), and dorsal (C) views.
Taxons Sauraechmodon

Dentary of Echinodon becklesii from the Lower Cretaceous Purbeck Formation of England. Left dentary (NHMUK 48215b) in lateral (A), medial (B), and dorsal (C) views.

Crétacé Echinodon Sauraechmodon formation
Pterodactylus compressirostris, holotype NHMUK PV 39410 (Cenomanian / Turonian, Chalk Formation). A–D proposed lectotype, fragment of the mandibular symphysis A left lateral view B respective line drawing C dorsal view D respective line drawing. E–H referred specimen, portion of the rostrum E left lateral view F respective line drawing G ventral view H respective line drawing. Abbreviations: ch – choanae, d – dentary, m – maxillae, naof – nasoantorbital fenestra, pl – palatine, pm – premaxillae, prid – palatal ridge, sul– sulcus. Arrows indicate alveoli or teeth. Scale bar = 10 mm. Photos courtesy of The Natural History Museum.
Taxons Lonchodectes

Pterodactylus compressirostris, holotype NHMUK PV 39410 (Cenomanian / Turonian, Chalk Formation). A–D proposed lectotype, fragment of the mandibular symphysis A left lateral view B respective line drawing C dorsal view D respective line drawing. E–H referred specimen, portion of the rostrum E left lateral view F respective line drawing G ventral view H respective line drawing. Abbreviations: ch – choanae, d – dentary, m – maxillae, naof – nasoantorbital fenestra, pl – palatine, pm – premaxillae, prid – palatal ridge, sul– sulcus. Arrows indicate alveoli or teeth. Scale bar = 10 mm. Photos courtesy of The Natural History Museum.

écaille dessin musée Cénomanien +8
Pterodactylus compressirostris, holotype NHMUK PV 39410 (Cenomanian / Turonian, Chalk Formation). A–D proposed lectotype, fragment of the mandibular symphysis A left lateral view B respective line drawing C dorsal view D respective line drawing. E–H referred specimen, portion of the rostrum E left lateral view F respective line drawing G ventral view H respective line drawing. Abbreviations: ch – choanae, d – dentary, m – maxillae, naof – nasoantorbital fenestra, pl – palatine, pm – premaxillae, prid – palatal ridge, sul– sulcus. Arrows indicate alveoli or teeth. Scale bar = 10 mm. Photos courtesy of The Natural History Museum.
Taxons Lonchodraconidae

Pterodactylus compressirostris, holotype NHMUK PV 39410 (Cenomanian / Turonian, Chalk Formation). A–D proposed lectotype, fragment of the mandibular symphysis A left lateral view B respective line drawing C dorsal view D respective line drawing. E–H referred specimen, portion of the rostrum E left lateral view F respective line drawing G ventral view H respective line drawing. Abbreviations: ch – choanae, d – dentary, m – maxillae, naof – nasoantorbital fenestra, pl – palatine, pm – premaxillae, prid – palatal ridge, sul– sulcus. Arrows indicate alveoli or teeth. Scale bar = 10 mm. Photos courtesy of The Natural History Museum.

écaille dessin musée Cénomanien +8
Pterodactylus compressirostris, holotype NHMUK PV 39410 (Cenomanian / Turonian, Chalk Formation). A–D proposed lectotype, fragment of the mandibular symphysis A left lateral view B respective line drawing C dorsal view D respective line drawing. E–H referred specimen, portion of the rostrum E left lateral view F respective line drawing G ventral view H respective line drawing. Abbreviations: ch – choanae, d – dentary, m – maxillae, naof – nasoantorbital fenestra, pl – palatine, pm – premaxillae, prid – palatal ridge, sul– sulcus. Arrows indicate alveoli or teeth. Scale bar = 10 mm. Photos courtesy of The Natural History Museum.
Taxons Lonchodectidae

Pterodactylus compressirostris, holotype NHMUK PV 39410 (Cenomanian / Turonian, Chalk Formation). A–D proposed lectotype, fragment of the mandibular symphysis A left lateral view B respective line drawing C dorsal view D respective line drawing. E–H referred specimen, portion of the rostrum E left lateral view F respective line drawing G ventral view H respective line drawing. Abbreviations: ch – choanae, d – dentary, m – maxillae, naof – nasoantorbital fenestra, pl – palatine, pm – premaxillae, prid – palatal ridge, sul– sulcus. Arrows indicate alveoli or teeth. Scale bar = 10 mm. Photos courtesy of The Natural History Museum.

écaille dessin musée Cénomanien +8
Holotype specimen of Hylaeosaurus armatus Mantell, 1833 (NMH R3775) from the Grinstead Clay Formation (late Valanginian) of Tilgate Forest, Sussex, southern England.
Taxons Hylaeosaurus

Holotype specimen of Hylaeosaurus armatus Mantell, 1833 (NMH R3775) from the Grinstead Clay Formation (late Valanginian) of Tilgate Forest, Sussex, southern England.

Valanginien holotype spécimen Hylaeosaurus +2
Holotype specimen of Hylaeosaurus armatus Mantell, 1833 (NMH R3775) from the Grinstead Clay Formation (late Valanginian) of Tilgate Forest, Sussex, southern England.
Taxons Hylosaurus

Holotype specimen of Hylaeosaurus armatus Mantell, 1833 (NMH R3775) from the Grinstead Clay Formation (late Valanginian) of Tilgate Forest, Sussex, southern England.

Valanginien holotype spécimen Hylaeosaurus +2
Figure description from the paper: "The segmented model of right lateral side of the skull of mixosaurid ichthyosaur Phalarodon fraasi (Merriam, 1910) PMO 235.393, from the Botneheia Formation, Middle Triassic of the Isfjorden area in Spitsbergen, Svalbard."
References:

Roberts, A.J.; Engelschiøn, V.S.; Hurum, J.H. (2022). "First three-dimensional skull of the Middle Triassic mixosaurid ichthyosaur Phalarodon fraasi from Svalbard, Norway". Acta Palaeontologica Polonica 67 (1): 51–62. DOI:https://doi.org/10.4202/app.00915.2021.
Taxons Phalarodon

Figure description from the paper: "The segmented model of right lateral side of the skull of mixosaurid ichthyosaur Phalarodon fraasi (Merriam, 1910) PMO 235.393, from the Botneheia Formation, Middle Triassic of the Isfjorden area in Spitsbergen, Svalbard." References: Roberts, A.J.; Engelschiøn, V.S.; Hurum, J.H. (2022). "First three-dimensional skull of the Middle Triassic mixosaurid ichthyosaur Phalarodon fraasi from Svalbard, Norway". Acta Palaeontologica Polonica 67 (1): 51–62. DOI:https://doi.org/10.4202/app.00915.2021.

description Norvège Trias moyen Trias +7
Figure description from the paper: "The segmented model of right lateral side of the skull of mixosaurid ichthyosaur Phalarodon fraasi (Merriam, 1910) PMO 235.393, from the Botneheia Formation, Middle Triassic of the Isfjorden area in Spitsbergen, Svalbard."
References:

Roberts, A.J.; Engelschiøn, V.S.; Hurum, J.H. (2022). "First three-dimensional skull of the Middle Triassic mixosaurid ichthyosaur Phalarodon fraasi from Svalbard, Norway". Acta Palaeontologica Polonica 67 (1): 51–62. DOI:https://doi.org/10.4202/app.00915.2021.
Taxons Contectopalatus

Figure description from the paper: "The segmented model of right lateral side of the skull of mixosaurid ichthyosaur Phalarodon fraasi (Merriam, 1910) PMO 235.393, from the Botneheia Formation, Middle Triassic of the Isfjorden area in Spitsbergen, Svalbard." References: Roberts, A.J.; Engelschiøn, V.S.; Hurum, J.H. (2022). "First three-dimensional skull of the Middle Triassic mixosaurid ichthyosaur Phalarodon fraasi from Svalbard, Norway". Acta Palaeontologica Polonica 67 (1): 51–62. DOI:https://doi.org/10.4202/app.00915.2021.

description Norvège Trias moyen Trias +7
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Actualités

Une ancienne sécheresse aurait pu anéantir les vrais hobbits il y a 61 000 ans
extinction formation
Une sécheresse massive qui a duré des siècles pourrait avoir entraîné l’extinction des « hobbits » de Flores. Les enregistrements climatiques conservés dans les formations souterraines montrent que les précipitations ont chuté au moment même où la petite espèce humaine disparaissait. Dans le même temps, le nombre d’éléphants pygmées dont ils dépendaient a fortement diminué à mesure que les rivières s’asséchaient. Avec la disparition de la nourriture et de l'eau, les hobbits ont peut-être été repoussés et entrés dans leur dernier chapitre.
19/02/2026 sciencedaily-human-evo ⚙ Traduction automatique
Épisode 171 : Mosasaures d'eau douce
Hell Creek Mosasaurus formation
Nous plongeons dans les eaux douces de la formation Hell Creek. Oh cher
12/12/2025 palaeocast ⚙ Traduction automatique
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