Trias

Intervalle géologique

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Figure 1. Evolution of macroecological traits in Dinosauria. Large scale event in dinosaur evolution (a); the origin of dinosaurs (star), hyperthermals (volcano), the earliest fossil Avialae (bird), the earliest fossil angiosperm (flower), the Cretaceous/Palaeogene mass extinction (asteroid). Phylogeny of dinosaurs (b) redrawn from Sereno and adapted to the current consensus and upon which an ancestral state reconstruction of temperature niche (mean annual temperature) after Chiarenza et al. is plotted; Mesozoic palaeogeographies (c) for Triassic (T), Jurassic (J) and Cretaceous (K). Silhouette colours symbolize body mass for each of the taxa represented; information on dietary habits are plotted after Barrett and Zanno & Makovicky; numbers represent clades discussed through this study: 1, Ornithischia; 2, Thyreophora; 3, Ornithopoda; 4, Hadrosauroidea; 5, Marginocephalia; 6, Ceratopsia; 7, Saurischia; 8, Sauropodomorpha; 9, Sauropoda; 10, Theropoda; 11, Ceratosauria; 12, Tetanurae; 13, Coelurosauria; 14, Maniraptoriformes; 15, Maniraptora; 16, Deinonychosauria; 17, Avialae; 18, Ornithothoraces. Palaeogeographies modified from original plots via R package ‘mapast’ using plate models by Scotese.

Figure 1. Evolution of macroecological traits in Dinosauria. Large scale event in dinosaur evolution (a); the origin of dinosaurs (star), hyperthermals (volcano), the earliest fossil Avialae (bird), the earliest fossil angiosperm (flower), the Cretaceous/Palaeogene mass extinction (asteroid). Phylogeny of dinosaurs (b) redrawn from Sereno and adapted to the current consensus and upon which an ancestral state reconstruction of temperature niche (mean annual temperature) after Chiarenza et al. is plotted; Mesozoic palaeogeographies (c) for Triassic (T), Jurassic (J) and Cretaceous (K). Silhouette colours symbolize body mass for each of the taxa represented; information on dietary habits are plotted after Barrett and Zanno & Makovicky; numbers represent clades discussed through this study: 1, Ornithischia; 2, Thyreophora; 3, Ornithopoda; 4, Hadrosauroidea; 5, Marginocephalia; 6, Ceratopsia; 7, Saurischia; 8, Sauropodomorpha; 9, Sauropoda; 10, Theropoda; 11, Ceratosauria; 12, Tetanurae; 13, Coelurosauria; 14, Maniraptoriformes; 15, Maniraptora; 16, Deinonychosauria; 17, Avialae; 18, Ornithothoraces. Palaeogeographies modified from original plots via R package ‘mapast’ using plate models by Scotese.

écaille Crétacé Jurassique Mésozoïque +23
Tawa is an early theropod from the Late Triassic. The genus is named after the Hopi word for the Puebloan sun god. It was a bipedal carnivore, estimated around 2.5 m in length, and weighing about 15 kg. A basal theropod, Tawa shares physical characteristics with coelophysoids and herrerasaurids, and its discovery supports the theory that dinosaurs originated in the southern supercontinent of Gondwanna, before diversifying as Pangea split apart.

Tawa is an early theropod from the Late Triassic. The genus is named after the Hopi word for the Puebloan sun god. It was a bipedal carnivore, estimated around 2.5 m in length, and weighing about 15 kg. A basal theropod, Tawa shares physical characteristics with coelophysoids and herrerasaurids, and its discovery supports the theory that dinosaurs originated in the southern supercontinent of Gondwanna, before diversifying as Pangea split apart.

Trias supérieur Trias Dinosauria Herrerasauridae +2
Various fossils pertaining to the holotype of the Triassic ichthyosaur Toretocnemus (originally Leptocheirus, also Merriamia) zitteli. This image is derived from plate 23 in Merriam (1903), done by an uncredited artist. The arrangement of the individual figures has been modified from the original (most notably, figures 3 and 4 were swapped to minimize confusion).
Original description:
Leptocheirus zitteli n. gen. and sp.
Figures reproduced natural size from the type specimen.

Fig. 1.— Right side of skull.
Fig. 2. — Cross-section of posterior portion of the lower jaw, taken above the point marked A on the lateral view of the skull.
Fig. 3. — Cross-section of the upper and lower jaws, taken at the break immediately behind the point marked Sp on the lateral view of the skull.
Fig. 4. — Lateral view of an anterior caudal centrum.
Legend (modified from original):

Ar - articular
A - angular
D - dentary
J - jugal
L - lacrimal
Mx - maxilla
Po - postorbital
Sa - surangular
Se - sclerotic ring
Sp - spenial
T - cross-section of tooth
X - doubtful element

Various fossils pertaining to the holotype of the Triassic ichthyosaur Toretocnemus (originally Leptocheirus, also Merriamia) zitteli. This image is derived from plate 23 in Merriam (1903), done by an uncredited artist. The arrangement of the individual figures has been modified from the original (most notably, figures 3 and 4 were swapped to minimize confusion). Original description: Leptocheirus zitteli n. gen. and sp. Figures reproduced natural size from the type specimen. Fig. 1.— Right side of skull. Fig. 2. — Cross-section of posterior portion of the lower jaw, taken above the point marked A on the lateral view of the skull. Fig. 3. — Cross-section of the upper and lower jaws, taken at the break immediately behind the point marked Sp on the lateral view of the skull. Fig. 4. — Lateral view of an anterior caudal centrum. Legend (modified from original): Ar - articular A - angular D - dentary J - jugal L - lacrimal Mx - maxilla Po - postorbital Sa - surangular Se - sclerotic ring Sp - spenial T - cross-section of tooth X - doubtful element

description Trias fossile holotype +7
The shoulder girdle of the holotype of the Triassic ichthyosaur Toretocnemus (originally Leptocheirus, also Merriamia) zitteli. This image is derived from plate 21 in Merriam (1903), done by an uncredited artist. The arrangement of the individual figures has been slightly modified from the original.
Original description: Leptocheirus zitteli n. gen. and sp.
Figures reproduced natural size from the type specimen.

Fig. 1.—Clavicles (cl) and probable interelavicle (Ic and Ic')- The interclavicle has been separated into two fragments by a crack in the matrix.
Fig. 2.—Inner side of the coracoids and the left scapula.

The shoulder girdle of the holotype of the Triassic ichthyosaur Toretocnemus (originally Leptocheirus, also Merriamia) zitteli. This image is derived from plate 21 in Merriam (1903), done by an uncredited artist. The arrangement of the individual figures has been slightly modified from the original. Original description: Leptocheirus zitteli n. gen. and sp. Figures reproduced natural size from the type specimen. Fig. 1.—Clavicles (cl) and probable interelavicle (Ic and Ic')- The interclavicle has been separated into two fragments by a crack in the matrix. Fig. 2.—Inner side of the coracoids and the left scapula.

description Trias holotype spécimen +5
Forelimb and partial hindlimb of the holotype of the Triassic ichthyosaur Toretocnemus (originally Leptocheirus, also Merriamia) zitteli. This image is derived from plate 22 in Merriam (1903), done by an uncredited artist.
Original description: Leptocheirus zitteli n. gen. and sp.
Figures reproduced natural size from the type specimen.

Fig. 1. — Superior side of left anterior limb. r, radius.
Fig. 2. — Posterior limb. t, tibia.

Forelimb and partial hindlimb of the holotype of the Triassic ichthyosaur Toretocnemus (originally Leptocheirus, also Merriamia) zitteli. This image is derived from plate 22 in Merriam (1903), done by an uncredited artist. Original description: Leptocheirus zitteli n. gen. and sp. Figures reproduced natural size from the type specimen. Fig. 1. — Superior side of left anterior limb. r, radius. Fig. 2. — Posterior limb. t, tibia.

membre description Trias holotype +7
Various fossils pertaining to the holotype of the Triassic ichthyosaur Toretocnemus californicus. This image is derived from plate 24 in Merriam (1903), done by an uncredited artist. The arrangement of the individual figures has been modified from the original.
Original description:
Toretocnemus californicus n. gen. and sp.
Figures reproduced natural size from the type specimen.

Fig. 1. — Inferior side of right posterior limb. t, tibia.
Fig. 2. — Right anterior limb. r, radius.
Fig. 3.— Pelvic arch.
Fig. 4. — Middle dorsal vertebrae and a rib from the same region.

Various fossils pertaining to the holotype of the Triassic ichthyosaur Toretocnemus californicus. This image is derived from plate 24 in Merriam (1903), done by an uncredited artist. The arrangement of the individual figures has been modified from the original. Original description: Toretocnemus californicus n. gen. and sp. Figures reproduced natural size from the type specimen. Fig. 1. — Inferior side of right posterior limb. t, tibia. Fig. 2. — Right anterior limb. r, radius. Fig. 3.— Pelvic arch. Fig. 4. — Middle dorsal vertebrae and a rib from the same region.

membre description Trias fossile +6
Illustration of some anterior caudal vertebrae of the Californian ichthyosaur Torteocnemus californicus (specimen UCMP 8100), from Merriam's 1908 monograph on American Triassic ichthyosaurs. This illustration appeared as figure 4 of plate 14; it has been modified to remove the stamp placed on top of it. While Merriam credits two illustrators as having produced artwork for this publication, it is not clear which of them created this image.

Illustration of some anterior caudal vertebrae of the Californian ichthyosaur Torteocnemus californicus (specimen UCMP 8100), from Merriam's 1908 monograph on American Triassic ichthyosaurs. This illustration appeared as figure 4 of plate 14; it has been modified to remove the stamp placed on top of it. While Merriam credits two illustrators as having produced artwork for this publication, it is not clear which of them created this image.

Trias spécimen Ichthyosauria Merriamia +1
Life restoration of the Triassic ichthyosaur Qianichthyosaurus xingyiensis.
References
Motani, Ryosuke; Jiang, Da-yong; Tintori, Andrea; Ji, Cheng; Huang, Jian-dong (2017). "Pre- versus post-mass extinction divergence of Mesozoic marine reptiles dictated by time-scale dependence of evolutionary rates". Proceedings of the Royal Society B 284 (1854): 20170241. DOI:10.1098/rspb.2017.0241.

Life restoration of the Triassic ichthyosaur Qianichthyosaurus xingyiensis. References Motani, Ryosuke; Jiang, Da-yong; Tintori, Andrea; Ji, Cheng; Huang, Jian-dong (2017). "Pre- versus post-mass extinction divergence of Mesozoic marine reptiles dictated by time-scale dependence of evolutionary rates". Proceedings of the Royal Society B 284 (1854): 20170241. DOI:10.1098/rspb.2017.0241.

écaille Mésozoïque Trias Ichthyosauria +3
Figure description from paper: "Figure 23: Early and Middle Triassic ichthyopterygian heads possessing longirostry.
Simplified outlines of four different Early and Middle Triassic ichthyopterygian heads possessing a long and slender rostrum. Specimens are at the same scale. (A) Utatsusaurus hataii (UHR 30691, Motani, Minoura & Ando, 1998); (B) Grippia longirostris (PMU R445, Motani, 2000); (C) Mixosaurus cornalianus (BES SC 1000, Renesto et al., 2020); (D) Besanosaurus leptorhynchus (PIMUZ T 4847, this paper); (E) Cymbospondylus buchseri (PIMUZ T 4351, Sander, 1989). Scale bar represents 10 cm."
References:

Bindellini, G.; Wolniewicz, A.S.; Miedema, F.; Scheyer, T.M.; Dal Sasso, C. (2021). "Cranial anatomy of Besanosaurus leptorhynchus Dal Sasso & Pinna, 1996 (Reptilia: Ichthyosauria) from the Middle Triassic Besano Formation of Monte San Giorgio, Italy/Switzerland: Taxonomic and palaeobiological implications". PeerJ 9: e11179. DOI:10.7717/peerj.11179.

Figure description from paper: "Figure 23: Early and Middle Triassic ichthyopterygian heads possessing longirostry. Simplified outlines of four different Early and Middle Triassic ichthyopterygian heads possessing a long and slender rostrum. Specimens are at the same scale. (A) Utatsusaurus hataii (UHR 30691, Motani, Minoura & Ando, 1998); (B) Grippia longirostris (PMU R445, Motani, 2000); (C) Mixosaurus cornalianus (BES SC 1000, Renesto et al., 2020); (D) Besanosaurus leptorhynchus (PIMUZ T 4847, this paper); (E) Cymbospondylus buchseri (PIMUZ T 4351, Sander, 1989). Scale bar represents 10 cm." References: Bindellini, G.; Wolniewicz, A.S.; Miedema, F.; Scheyer, T.M.; Dal Sasso, C. (2021). "Cranial anatomy of Besanosaurus leptorhynchus Dal Sasso & Pinna, 1996 (Reptilia: Ichthyosauria) from the Middle Triassic Besano Formation of Monte San Giorgio, Italy/Switzerland: Taxonomic and palaeobiological implications". PeerJ 9: e11179. DOI:10.7717/peerj.11179.

écaille description Italie Suisse +12
Besanosaurus leptorhynchus referred specimen PIMUZ T 4376, preserved on the same slab as a specimen of the smaller Mixosaurus. Scale bar represents 50 cm (1.6 ft). Figure 3C of Bindellini et al. (2021).
References:

Bindellini, G.; Wolniewicz, A.S.; Miedema, F.; Scheyer, T.M.; Dal Sasso, C. (2021). "Cranial anatomy of Besanosaurus leptorhynchus Dal Sasso & Pinna, 1996 (Reptilia: Ichthyosauria) from the Middle Triassic Besano Formation of Monte San Giorgio, Italy/Switzerland: Taxonomic and palaeobiological implications". PeerJ 9: e11179. DOI:10.7717/peerj.11179.

Besanosaurus leptorhynchus referred specimen PIMUZ T 4376, preserved on the same slab as a specimen of the smaller Mixosaurus. Scale bar represents 50 cm (1.6 ft). Figure 3C of Bindellini et al. (2021). References: Bindellini, G.; Wolniewicz, A.S.; Miedema, F.; Scheyer, T.M.; Dal Sasso, C. (2021). "Cranial anatomy of Besanosaurus leptorhynchus Dal Sasso & Pinna, 1996 (Reptilia: Ichthyosauria) from the Middle Triassic Besano Formation of Monte San Giorgio, Italy/Switzerland: Taxonomic and palaeobiological implications". PeerJ 9: e11179. DOI:10.7717/peerj.11179.

écaille Italie Suisse Trias moyen +8
Modified version of figure 23 of Bindellini et al. (2021) showing just the ichthyosaurs from the Besano Formation: Besanosaurus leptorhynchus (A), Mixosaurus cornalianus (B), and Cymbospondylus buchseri (C). The original figure was uploaded unmodified here: File:Early to Middle Triassic Longirostrine Ichthyopterygians.png
References:

Bindellini, G.; Wolniewicz, A.S.; Miedema, F.; Scheyer, T.M.; Dal Sasso, C. (2021). "Cranial anatomy of Besanosaurus leptorhynchus Dal Sasso & Pinna, 1996 (Reptilia: Ichthyosauria) from the Middle Triassic Besano Formation of Monte San Giorgio, Italy/Switzerland: Taxonomic and palaeobiological implications". PeerJ 9: e11179. DOI:10.7717/peerj.11179.

Modified version of figure 23 of Bindellini et al. (2021) showing just the ichthyosaurs from the Besano Formation: Besanosaurus leptorhynchus (A), Mixosaurus cornalianus (B), and Cymbospondylus buchseri (C). The original figure was uploaded unmodified here: File:Early to Middle Triassic Longirostrine Ichthyopterygians.png References: Bindellini, G.; Wolniewicz, A.S.; Miedema, F.; Scheyer, T.M.; Dal Sasso, C. (2021). "Cranial anatomy of Besanosaurus leptorhynchus Dal Sasso & Pinna, 1996 (Reptilia: Ichthyosauria) from the Middle Triassic Besano Formation of Monte San Giorgio, Italy/Switzerland: Taxonomic and palaeobiological implications". PeerJ 9: e11179. DOI:10.7717/peerj.11179.

Italie Suisse Trias moyen Trias +7
Utatsusaurus hataii, an Early Triassic Ichthyosaur from Japan, pencil drawing
Taxons Grippiidae

Utatsusaurus hataii, an Early Triassic Ichthyosaur from Japan, pencil drawing

dessin Japon Early Triassic Trias +4
Augustasaurus hagdorni, a pistosaur from the Middle Triassic of nevada. Digital.
Taxons Pistosauroidea

Augustasaurus hagdorni, a pistosaur from the Middle Triassic of nevada. Digital.

Trias moyen Trias Augustasaurus Pistosauria +2
Augustasaurus hagdorni, a pistosaur from the Middle Triassic of nevada. Digital.
Taxons Pistosauria

Augustasaurus hagdorni, a pistosaur from the Middle Triassic of nevada. Digital.

Trias moyen Trias Augustasaurus Pistosauria +2
Nothosaurus mirabilis, a Nothosaur from the Triassic of Europe, North Africa and Russia, pencil drawing
Taxons Nothosauria

Nothosaurus mirabilis, a Nothosaur from the Triassic of Europe, North Africa and Russia, pencil drawing

dessin Russie Trias Nothosauria +2
Nothosaurus mirabilis, a Nothosaur from the Triassic of Europe, North Africa and Russia, pencil drawing
Taxons Nothosauroidea

Nothosaurus mirabilis, a Nothosaur from the Triassic of Europe, North Africa and Russia, pencil drawing

dessin Russie Trias Nothosauria +2
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Actualités

Un nouveau dinosaure carnivore du Trias identifié à Ghost Ranch
États-Unis Trias supérieur Trias fossile spécimen Coelophysis Dinosauria Ptychotherates nouvelle espèce crâne
Des chercheurs de Virginia Tech (Virginie, États-Unis) ont identifié une nouvelle espèce de dinosaure du Trias supérieur.  Le dinosaure a été nommé Ptychotherates bucculentus.  Ce nouveau taxon de dinosaure carnivore a été érigé sur la base de l'examen détaillé d'un fossile de crâne incomplet, bien conservé mais mélangé (numéro de spécimen CM 31368).  Le crâne provient de la Coelophysis de renommée mondiale
11/05/2026 everythingdinosaur ⚙ Traduction automatique
New Fossil from Brazil Reveals Unexpected Diversity among Pre-Dinosaur Herbivores
Un nouveau fossile du Brésil révèle une diversité inattendue parmi les herbivores pré-dinosaures
mâchoire Brésil Trias fossile Dinosauria nouvelle espèce partiel crâne
Des paléontologues du Brésil ont identifié un nouveau genre et une nouvelle espèce de rhynchosaure hyperodapédontique – une lignée éteinte de reptiles herbivores à bec – sur la base d'un crâne partiel et de mâchoires inférieures récupérés dans des roches du Trias. L'article Un nouveau fossile du Brésil révèle une diversité inattendue parmi les herbivores pré-dinosaures est apparu en premier sur Sci.News : Breaking Science News.
23/04/2026 sci-news ⚙ Traduction automatique
Museum Fossil Reveals Triassic Crocodile Cousin with Powerful Jaws
Un fossile de musée révèle un cousin crocodile du Trias doté de mâchoires puissantes
mâchoire musée Trias supérieur Trias fossile spécimen tomographie nouvelle espèce
Les tomodensitogrammes d'un spécimen vieux de plusieurs décennies du Musée d'histoire naturelle de Yale Peabody montrent une nouvelle espèce de crocodylomorphe au museau court et aux mâchoires inhabituellement fortes, offrant un instantané rare de la spécialisation écologique à l'époque du Trias supérieur. Le post Museum Fossil révèle un cousin crocodile du Trias doté de mâchoires puissantes est apparu en premier sur Sci.News: Breaking Science News.
20/04/2026 sci-news ⚙ Traduction automatique
New Triassic Dinosaur Species Identified in New Mexico
De nouvelles espèces de dinosaures du Trias identifiées au Nouveau-Mexique
Mexique États-Unis Trias Dinosauria nouvelle espèce crâne
Un nouveau genre et une nouvelle espèce de dinosaure herrerasaurien carnivore ont été décrits à partir d'un crâne incomplet mais bien conservé trouvé dans le nord du Nouveau-Mexique, aux États-Unis. L'article Nouvelles espèces de dinosaures du Trias identifiées au Nouveau-Mexique est apparu en premier sur Sci.News : Breaking Science News.
17/04/2026 sci-news ⚙ Traduction automatique
Un fossile écrasé révèle un dinosaure qui n’aurait pas dû exister
Trias fossile Dinosauria découverte extinction nouvelle espèce crâne
Un crâne de dinosaure gravement mutilé, oublié dans un tiroir, s'est avéré être une découverte rare et importante. Reconstruit par un étudiant de Virginia Tech, il a révélé une nouvelle espèce de dinosaure carnivore primitif présentant des caractéristiques inhabituelles jamais vues auparavant. Le fossile suggère que certains groupes de dinosaures ont été anéantis lors de l’extinction de la fin du Trias, et pas seulement leurs rivaux. Il s’agit peut-être de l’un des derniers survivants d’une ancienne lignée de dinosaures.
15/04/2026 sciencedaily ⚙ Traduction automatique
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