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Ceratopsipes goldenensis Lockley & Hunt, 1995 - ceratopsian dinosaur trackway in the Cretaceous of Colorado, USA. Ceratopsians are the "horned dinosaurs". They were large, quadrupedal, herbivorous dinosaurs having a beaked skull and a frill - an extension of bone behind the skull that partially covered the neck. Ceratopsian dinosaurs are known from the Jurassic and Cretaceous. The last members of the group died out at the Cretaceous-Tertiary boundary, 65 million years ago. Seen here is a ceratopsian dinosaur trackway. Ceratopsian footprints are very rare. The best examples are in Colorado's Laramie Formation, a nonmarine, coastal plain to deltaic succession of mostly mixed siliciclastic sedimentary rocks - sandstones, siltstones, claystones, and coals. The footprints here consist of convex bulges on the basal surfaces of structurally-tilted sandstone beds. The ichnospecies Ceratopsipes goldenensis was named based on footprints at this locality. The trackmaker was very likely Triceratops, the # 1 most famous ceratopsian dinosaur. From on-site signage: Triceratops Tracks Several tracks of Triceratops, or a closely related horned dinosaur were first discovered in this area and named Ceratopsipes goldenensis (meaning tracks of a horned dinosaur from Golden). By happy coincidence, the first Triceratops known to science also comes from the greater Denver area near 13th Avenue and Federal Blvd. Growing to a length of almost 25 feet, Triceratops and other members of the horned dinosaur family were herbivorous animals that roamed in herds. Their trademark horns and frills that covered their necks probably served for defense. A replica Triceratops skull is on display in the clubhouse at the Fossil Trace Golf Club. These tracks (identified by small signs) help us understand that Triceratops walked with a narrow gait and erect front limbs than with a wide, sprawling gait as previously depicted. In 1887, the first Triceratops bones known to science were also discovered locally at a site near 13th Avenue and Federal Boulevard in Denver. Nearby one may see many other traces of life such as burrows, probably made by invertebrates and impressions of plant debris. Note that you are looking at all tracks and traces from the underside - in negative aspect - what one might call a worm’s eye view. All of these trace fossils indicate that the Golden area once had a warm, temperate to subtropical climate. Classification: Animalia, Chordata, Vertebrata, Reptilia, Archosauria, Dinosauria, Ornithischia, Marginocephalia, Ceratopsia, Ceratopsidae Stratigraphy: Laramie Formation, Maastrichtian Stage, upper Upper Cretaceous Locality: outcrop along Triceratops Trail, Parfet Prehistoric Preserve, southern side of the town of Golden, Colorado, USA (~vicinity of 39° 44' 35.24" North latitude, 105° 13’ 09.69" West longitude) Some info. from: Lockley & Hunt (1995) - Ceratopsid tracks and associated ichnofauna from the Laramie Formation (Upper Cretaceous: Maastrichtian) of Colorado. Journal of Vertebrate Paleontology 15: 592-614.

Skeletal diagram featuring the optimal remains of the holotype of Graciliceratops mongoliensis: ZPAL MgD-I/156.[1] Found in the Bayan Shireh Formation, in the original description the remains were referred to the genus Microceratops (now obsolete).[2] However, Sereno in 2000 noted that there was no base for this referral, then, he created a new genus and species for this specimen.[1] The holotype is very fragmented (specially the skull), consisting of:[2] Fragmented skull; 4 cervical, 12 dorsal and 7 sacral vertebrae; right scapula; proximal end of left scapula; left coracoid; right humerus, radius and fragmentary ulna; proximal and distal end of left humerus; proximal fragments of both pubis; fragments of both illium and fragment of right ischium; right femur, tibia and nearly complete pes; distal part of left tibia, fragmentary left pes; tarsals and isolated ribs.[2] The sacral vertebrae are not fused, an indicator of the immaturity of this specimen; the estimated adult size is about 2 meters long or similar to Protoceratops.[2][1] Right quadratojugal, quadrate and fragmentary jugal were reversed in order to get an optimal view.

Braincase of WSC 10058, holotype of Ornatops Braincase of Ornatops incantatus in digital 3D model in right lateral (A and C) and left lateral (D and F) views, with enlarged images of the right (B) and left (E) auditory regions. Credit: McDonald AT, Wolfe DG, Freedman Fowler EA, Gates TA. 2021. A new brachylophosaurin (Dinosauria: Hadrosauridae) from the Upper Cretaceous Menefee Formation of New Mexico. PeerJ 9:e11084 https://doi.org/10.7717/peerj.11084

The hadrosaurid dinosaur Wulagasaurus dongi gen. et sp. nov. from the Upper Cretaceous Yuliangze Formation at the Wulaga quarry, China. A. Holotype GMH W184, right dentary in lateral (A1) and medial (A2) views. B. GMH W217, right dentary in lateral (B1) and medial (B2) views.

Figure 1: Reconstruction of the skull of Bellusaurus sui from the Middle-Late Jurassic Shishugou Formation of Xinjiang, China. This reconstruction is a composite based on isolated holotypic and referred material. (A) Right lateral view. (B) Dorsal view. Holotypic elements are indicated in blue and referred elements are in green.

Cervical vertebrae of rebbachisaurid sauropod Katepensaurus goicoecheai Ibiricu, Casal, Martínez, Lamanna, Luna, and Salgado, 2013a from the Cenomanian–Turonian Bajo Barreal Formation of Chubut Province, Argentina. A. UNPSJB-PV 1007/1, anterior cervical vertebra in anterior (A1) and right ventrolateral (A2) views. B. UNPSJB-PV 1007/2, anterior cervical vertebra in right lateral view. C. UNPSJB-PV 1007/3, middle cervical vertebra in right lateral view. Lateral fossae of the centrum (hypothesized as pneumatic in origin) indicated by arrows.

Reconstruction of the holotype material of Silutitan sinensis (IVPP V27874), a Cretaceous euhelopid sauropod from the Chinese Shengjinkou Formation, scaled after Euhelopus zdanskyi. Silhouette modified from "Euhelopus zdanskyi Skeletal" by Gunnar Bivens (CC BY 3.0) (https://www.deviantart.com/gunnarbivens/art/Euhelopus-zdanskyi-Skeletal-833724646)

This file represents a possible life appearance of the Indian Titanosaurian Sauropod dinosaur Jainosaurus septentrionalis from the Late Cretaceous (Maastrichtian) of India, belonging to the Lameta Formation. References used: Huene & Matley (1933) Hunt et al. (1995) Gunnar Bivens' skeletal diagram Wilson et al. (2008) "Reassessment of Sauropod Dinosaur Jainosaurus (="Antarctosaurus") Septentrionalis from the Upper Cretaceous of India"

Representative taxa from the early Campanian Muthmannsdorf fauna from the Grünbach Formation, eastern Austria. A Doratodon carcharidens (Mesoeucrocodylia) mandible (PIUW 2349/57) in dorsal view (photo by Márton Rabi) B Indeterminate azhdarchid (Pterosauria, Azhdarchidae), left humerus (PIUW 2349/102) in anterior view C ‘Megalosaurus pannoniensis’ basal tetanuran (Theropoda, Tetanurae), tooth (PIUW uncatalogued) in lateral view D Mochlodon suessi (Ornithopoda, Rhabdodontidae), right dentary (holotype, PIUW 2349/2) in medial view. Scale bars equal 2 cm in A, B and D and 1 cm in C.

Right dentary of Zalmoxes shqiperorum holotype NHMUK R4900in dorsal and lateral views, from the Sinpetru Formation of Sânpetru (Romania; outline based on the specimen UBB NVZ1-1).

Dentaries of selected European rhabdodontomorphs in dorsal and lateral views. (A) Obelignathus septimanicus, holotype MDE D30, right dentary from the 'Grès à Reptiles' Formation, Montouliers (France); (B) Mochlodon vorosi holotype MTM V 2010.105.1, left dentary from the Csehbánya Formation, Iharkút (Hungary; mirrored); (C) Mochlodon suessi, lectotype PIUW 2349/2, right dentary from the Grünbach Formation, Muthmannsdorf (Austria); (D) Zalmoxes robustus holotype NHMUK R3392, right dentary from the Sinpetru Formation, Sânpetru (Romania); (E) Rhabdodon priscus, lectotype MPLM 30, left dentary from the Marnes Rouges Inférieures Formation, la Nerthe (France; mirrored drawing after Matheron; outline based on specimens MPLM 30 and MPLM 31, size estimation based on Matheron and Brinkman); (F) Zalmoxes shqiperorum holotype NHMUK R4900, right dentary from the Sinpetru Formation of Sânpetru (Romania; outline based on the specimen UBB NVZ1-1).

Representative taxa from the Santonian Iharkút fauna from the Csehbánya Formation, Bakony Mountains, western Hungary. A Pannoniasaurus inexpectatus (Squamata, Mosasauroidea), dorsal vertebra (MTM uncatalogued) in dorsal view (photo by Réka Kalmár) B Foxemys trabanti (Pleurodira, Bothremydidae), skull (MTM V 2010.215.1.) in dorsal view (photo by Márton Rabi). C Bicuspidon aff. hatzegiensis (Squamata, Borioteiioidea), left dentary (MTM 2006.112.1.) in medial view (photo by László Makádi) D Basal tetanuran (Theropoda, Tetanurae), tooth (MTM V.01.54) in ?lingual view E Indeterminate abelisaurid (Theropoda, Abelisauridae), pedal ungual phalanx (MTM V 2008.43.1.) in lateral view F Pneumatoraptor fodori (Theropoda, Paraves), left scapulocoracoid (holotype, MTM V 2008.38.1.) in lateral view G Mochlodon vorosi (Ornithopoda, Rhabdodontidae), left dentary (holotype, MTM V 2010.105.1) in lateral view H Bakonydraco galaczi (Pterosauria, Azhdarchidae), mandible (holotype, MTM 2007.110.1) in dorsal view I Iharkutosuchus makadii (Eusuchia, Hylaeochampsidae), skull (holotype, MTM 2006.52.1) in dorsal view J Hungarosaurus tormai (Ankylosauria, Nodosauridae), right dentary (MTM 2007.25.2) in lateral view K Bauxitornis mindszentyae (Aves, Enantiornithes), left tarsometatarsus (holotype, MTM V 2009.38.1) in anterior view L Ajkaceratops kozmai (Ceratopsia), fused rostral and premaxillae (holotype, MTM V 2009.192.1) in lateral view. Scale bars: 2 cm in A, V, G, H, I, J; 1 cm in D, E, F, K, L; 1 mm in C.

Representative taxa from the latest Campanian–Maastrichtian faunas from Transylvania, western Romania. A–B Nidophis insularis (Serpentes, Madtsoiidae), articulated vertebrae (LPB (FGGUB) v.547/2) in left lateral (A) and dorsal (B) views (Densuş-Ciula Formation, Tuştea, Haţeg Basin; photo by Ştefan Vasile) C Allodaposuchus precedens (Eusuchia, ?Hylaeochampsidae), skull (PSMUBB V 438) in dorsal view (Sebeş = Şard Formation, Oarda de Jos, southwestern Transylvanian Basin; photo by Vlad Codrea/Massimo Delfino) D Theriosuchus sympiestodon (Mesoeucrocodylia, Atoposauridae), right maxilla (MCDRD 793) in lateral view (Sînpetru Formation, Sînpetru, Haţeg Basin) E–F Indeterminate titanosaur (?Magyarosaurus dacus) (Sauropoda, Titanosauria), isolated osteoderm (LPB (FGGUB) R.1410) in dorsal (E) and lateral (F) views (Sînpetru Formation, Sînpetru, Haţeg Basin) G Indeterminate ornithuran bird (Aves, Ornithurae), incomplete left tibiotarsus (LPB (FGGUB) R.1902) in anterior view (Densuş-Ciula Formation, Vălioara, Haţeg Basin) H Balaur bondoc (Theropoda, Dromaeosauridae), articulated left distal hindlimb (EME PV.313) in lateral view (Sebeş = Şard Formation, Sebeş-Glod, southewestern Transylvanian Basin; photo by Mick Ellison) I Zalmoxes robustus (Ornithopoda, Rhabdodontidae), right dentary (NHMUK R.3407) in medial view (Sînpetru Formation, Sînpetru, Haţeg Basin) J Telmatosaurus transsylvanicus (Hadrosauria), right maxilla (MFGI unnumbered) in lateral view (Sînpetru Formation, Sînpetru, Haţeg Basin) K Indeterminate nodosaurid – Struthiosaurus transylvanicus or new taxon – (Ankylosauria, Nodosauridae), isolated tooth (LPB (FGGUB) R.2182) in medial view (Sînpetru Formation, Sînpetru, Haţeg Basin) L Barbatodon transylvanicus (Multituberculata, Kogaionidae), right maxilla (LPB (FGGUB) M.1635) in medial view (Sînpetru Formation, Pui, Haţeg Basin). Scale bars equal 1 mm in A, B; 5 mm in K; 1 cm in G, L; 2 cm in D; and 5 cm in C, E, F, H, I, J.

Palaeogeographic distribution of late Early and early Late Cretaceous pterosaur assemblages. Taxonomic composition of assemblages shown on Fig. 1. Palaeogeography based on Smith et al. 1994. Abbreviations: 1. Cambridge Greensand, England: 2. Lower Chalk, England: 3. Züümbayan Svita, Khuren-Dukh, Mongolia: 4. Lysaya Gora, Saratov, Russia: 5. Kem Kem red beds, Morocco: 6. Paw Paw Formation, Texas, USA: 7. Lagarcito Formation, San Luis, Argentina: 8. Santana and Crato Formations, Ceara, Brazil: 9. Toolebuc Formation, Queensland, Australia.

Locality map for Australian eurypodan thyreophoran fossils. 1, Stegosaurian? footprint (QM F5701), Walloon Coal Measures, Balgowan Colliery, Balgowan (Bajocian–Bathonian); 2, Minmi paravertebra holotype (QM F10329) (Molnar, 1980), Minmi Member, Bungil Formation (Valanginian–Barremian); 3, Thyreophoran trackways, Broome Sandstone, Dampier Peninsula, Western Australia (Valanginian–Barremian); 4, Ankylosauria indet. (see Barrett et al., 2010) ‘Flat Rocks’ Wonthaggi Formation (upper Hauterivian–Albian); 5, NMV P216739, ‘Lake Copco–Dinosaur Cove’ Eumeralla Formation (middle upper Aptian to lower middle Albian) (Barrett et al., 2010); 6, QM F33286; 7, AM F119849 and AM F35259; 8, Kunbarrasaurus ieversi gen. et sp. nov. (formerly Minmi sp.) (QM F18101); 9, QM F33565 and QM F33566; 10, QM F44324-28. Legend: Dark Green, Toolebuc Formation (late middle–early late Albian); Green, Allaru Formation (upper Albian–(?)lower Cenomanian); Light green, Mackunda Formation (upper Albian–lower Cenomanian); Lightest green, Winton Formation (late Albian–early Turonian).

Original figure caption: .mw-parser-output .smallcaps{font-variant:small-caps}The Middletown Slab covered with the Footprints of Carnivorous Dinosaurs. The tracks are in high relief. Additional notes: Most if not all of these tridactylous (i.e. three-toed) footprints/tracks (but not the actual trackmaker!) are referred to as Grallator or as Grallator-type trace fossils. “High relief” means that these are actually casts of footprints forming a positive relief on the lower surface of the sandstone slab (so-called positive hyporelief). The material that originally formed the mud over which the dinosaurs walked was too friable to be recovered from the quarry in one piece. The slab consists of so called ‘brownstone’ which is the trading name of the sandstone quarried at Middletown, Connecticut. This sandstone belongs to the Lower Jurassic Portland Formation of the Hartford Basin (“Connecticut Valley”) and thus to the upper part of the Newark Supergroup. The trackmakers probably were relatively small ‘primitive’ theropod dinosaurs (coelophysoids) such as Podokesaurus the remains of which were recovered from Lower Jurassic deposits of the Hartford Basin.