formation

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Galerie d'images

Drawings of Dryolestida left upper−molar, modified from the original descriptions. A–C, Kimmeridgian; D–F, Tithonian–Berriasian; G–H, early Barremian; I, late Barremian. A. Left M4 or M5 of Comotherium richi Prothero, 1981, from the Upper Jurassic Morrison Formation, Como Bluff locality, Wyoming, USA. B. Right M5, reversed, of Dryolestes leiriensis Martin, 1999, from Guimarota, Portugal.C. Left M6 of Krebsotherium lusitanicum Martin, 1999, from Guimarota, Portugal. D. Right M4 or M 6 (reversed) of Portopinheirodon asymmetricus Martin, 1999, Porto Pinheiro, Lourinha, Portugal. E. Right M5 (reversed) of Laolestes andresi Martin, 1999, from Porto Pinheiro, Lourinha, Portugal. F. Right upper molar of Donodon perscriptoris Sigogneau−Russell, 1991, Anoual, Morocco.G. Left M4 orM5 of Crusafontia amoae sp. nov., holotype, Cuesta Corrales 2, El Castellar Formation, Galve, Teruel, Spain. H. Right M6 orM7 (reversed) of Crusafontia amoae sp. nov., first described as upper molar of Crusafontia cuencana by Krebs (1993), P−2 H4 Pelejón 2, Galve, Teruel, Spain. I. Left M2 or M3, of Crusafontia cuencana Henkel and Krebs, 1969, Uña, La Huérguina Formation, Cuenca, Spain, (from Krebs 1993). Scale bars 1 mm.

Drawings of Dryolestida left upper−molar, modified from the original descriptions. A–C, Kimmeridgian; D–F, Tithonian–Berriasian; G–H, early Barremian; I, late Barremian. A. Left M4 or M5 of Comotherium richi Prothero, 1981, from the Upper Jurassic Morrison Formation, Como Bluff locality, Wyoming, USA. B. Right M5, reversed, of Dryolestes leiriensis Martin, 1999, from Guimarota, Portugal.C. Left M6 of Krebsotherium lusitanicum Martin, 1999, from Guimarota, Portugal. D. Right M4 or M 6 (reversed) of Portopinheirodon asymmetricus Martin, 1999, Porto Pinheiro, Lourinha, Portugal. E. Right M5 (reversed) of Laolestes andresi Martin, 1999, from Porto Pinheiro, Lourinha, Portugal. F. Right upper molar of Donodon perscriptoris Sigogneau−Russell, 1991, Anoual, Morocco.G. Left M4 orM5 of Crusafontia amoae sp. nov., holotype, Cuesta Corrales 2, El Castellar Formation, Galve, Teruel, Spain. H. Right M6 orM7 (reversed) of Crusafontia amoae sp. nov., first described as upper molar of Crusafontia cuencana by Krebs (1993), P−2 H4 Pelejón 2, Galve, Teruel, Spain. I. Left M2 or M3, of Crusafontia cuencana Henkel and Krebs, 1969, Uña, La Huérguina Formation, Cuenca, Spain, (from Krebs 1993). Scale bars 1 mm.

description dessin Maroc Portugal +11
Holotype specimen TMP 2000.29.01 of the ophthalmosaurian ichthyosaur Athabascasaurus bitumineus from the Lower Cretaceous Clearwater Formation of Alberta, in Royal Tyrrell Museum, Drumheller, Alberta, Canada.

Holotype specimen TMP 2000.29.01 of the ophthalmosaurian ichthyosaur Athabascasaurus bitumineus from the Lower Cretaceous Clearwater Formation of Alberta, in Royal Tyrrell Museum, Drumheller, Alberta, Canada.

musée Canada Crétacé holotype +5
Fig 2. Chrono-, bio- and lithostratigraphic context of the Reuchenette Formation in the Ajoie district, Canton Jura, NW Switzerland (modified from [20,23,28,31]. Four track-bearing intervals, named lower, intermediate, and upper (dinosaur track) levels, and track levels 600 have been identified within the Courtedoux Member (Nerinean Limestones, sensu [32]). All studied material comes from the intermediate (levels 1000–1100) and upper (levels 1500–1650) dinosaur track levels, details shown on inset on the upper right.

Fig 2. Chrono-, bio- and lithostratigraphic context of the Reuchenette Formation in the Ajoie district, Canton Jura, NW Switzerland (modified from [20,23,28,31]. Four track-bearing intervals, named lower, intermediate, and upper (dinosaur track) levels, and track levels 600 have been identified within the Courtedoux Member (Nerinean Limestones, sensu [32]). All studied material comes from the intermediate (levels 1000–1100) and upper (levels 1500–1650) dinosaur track levels, details shown on inset on the upper right.

Suisse Reuchenette Dinosauria Megalosauripus +1
Remake of the original picture of the Drzewica Formation. Terrestrial environment of the Pliensbachian-Toarcian boundary of Fennoscandinavia  Inland environment of the  Bornholm Coast, nearby the German realm of the Ciechocinek Formation. Includes
Ciechocinek Formation (Lower Toarcian, Bones) and Drzwica Formation (Latest Pliensbachian, Footprints) Fauna
Sorthat Formation environment, fluvial influenced mainland with Cheirolepidaceae and Bennetitales as dominant flora
Dinosaurs are based on material found on various locations of Northern Germany, and Footprints of the underliying Drzewica Formation at the Holy Cross Mountains, connected with Bornholm at the time.
Dinosaur Species appeared: 

Megalosauripus isp. Large Footprints (+65 cm) found on the Drzewica Formation. There is a dorsal vertebrae on the German Margin of the Ciechocinek Formation assigend to Megalosauria (Huene, 1966).
Gravisauria spp. representing the Grimmen Sauropod reported on 2014, as a taxon related with Tazoudasaurus. Barapasaurus-like footprints are know from the Drzewica Formation.
Coelophysoidea spp. based on coeval Anchisauripus tracks from the Holy Cross Mountains.
Basal Ornithischan, related to Eocursor, based on a crouching trace (Gerard Dariusz Gierlinski, Martin G. Lockley, Grzegorz Niedźwiedzki:2009).
Massospondylidae spp. based on Otozum-like tracks.
Taxons Megalosauripus

Remake of the original picture of the Drzewica Formation. Terrestrial environment of the Pliensbachian-Toarcian boundary of Fennoscandinavia Inland environment of the Bornholm Coast, nearby the German realm of the Ciechocinek Formation. Includes Ciechocinek Formation (Lower Toarcian, Bones) and Drzwica Formation (Latest Pliensbachian, Footprints) Fauna Sorthat Formation environment, fluvial influenced mainland with Cheirolepidaceae and Bennetitales as dominant flora Dinosaurs are based on material found on various locations of Northern Germany, and Footprints of the underliying Drzewica Formation at the Holy Cross Mountains, connected with Bornholm at the time. Dinosaur Species appeared: Megalosauripus isp. Large Footprints (+65 cm) found on the Drzewica Formation. There is a dorsal vertebrae on the German Margin of the Ciechocinek Formation assigend to Megalosauria (Huene, 1966). Gravisauria spp. representing the Grimmen Sauropod reported on 2014, as a taxon related with Tazoudasaurus. Barapasaurus-like footprints are know from the Drzewica Formation. Coelophysoidea spp. based on coeval Anchisauripus tracks from the Holy Cross Mountains. Basal Ornithischan, related to Eocursor, based on a crouching trace (Gerard Dariusz Gierlinski, Martin G. Lockley, Grzegorz Niedźwiedzki:2009). Massospondylidae spp. based on Otozum-like tracks.

os Allemagne Pliensbachien Toarcien +7
Left ilium of the camarasauromorph sauropod Brontomerus mcintoshi from the Lower Cretaceous Cedar Mountain Formation of Utah, type specimen OMNH 66430 in lateral view reconstructed from the three fragments (A), and ventral view (B).
Taxons Brontomerus

Left ilium of the camarasauromorph sauropod Brontomerus mcintoshi from the Lower Cretaceous Cedar Mountain Formation of Utah, type specimen OMNH 66430 in lateral view reconstructed from the three fragments (A), and ventral view (B).

Cedar Mountain Crétacé spécimen Brontomerus +1
Right hind limb of the sauropodomorph dinosaur Musankwa sanyatiensis gen. et sp. nov. (NHMZ 2521) from the Pebbly Arkose Formation (Norian, Upper Triassic) of Spurwing Island, Zimbabwe. A. Right femur in posterior (A1), lateral (A2), anterior (A3), medial (A4), proximal (A5), and distal (A6) views. B. Right tibia with conjoined astragalus in anterior (B1), lateral (B2), posterior (B3), medial (B4), and proximal (B5) views.
Taxons Musankwa

Right hind limb of the sauropodomorph dinosaur Musankwa sanyatiensis gen. et sp. nov. (NHMZ 2521) from the Pebbly Arkose Formation (Norian, Upper Triassic) of Spurwing Island, Zimbabwe. A. Right femur in posterior (A1), lateral (A2), anterior (A3), medial (A4), proximal (A5), and distal (A6) views. B. Right tibia with conjoined astragalus in anterior (B1), lateral (B2), posterior (B3), medial (B4), and proximal (B5) views.

membre Zimbabwe Norien Trias +3
Lonchodraco giganteus comb. n. Lectotype NHMUK PV 39412 (Cenomanian / Turonian, Chalk Formation). A–F articulated anterior parts of the rostrum and mandible A right lateral view B respective line drawing C left lateral view D respective line drawing E anterior view F respective line drawing G associated scapulocoracoid in posterior view. Abbreviations: ch – choanae, cor – coracoid, d – dentary, dcr – dentary crest, m – maxillae, pl – palatine, pm – premaxillae, pmcr – premaxillaery crest, prid – palatal ridge, sca – scapula. Arrows indicate alveoli or teeth. Scale bar = 10 mm. Photos courtesy of The Natural History Museum.
Taxons Lonchodraco

Lonchodraco giganteus comb. n. Lectotype NHMUK PV 39412 (Cenomanian / Turonian, Chalk Formation). A–F articulated anterior parts of the rostrum and mandible A right lateral view B respective line drawing C left lateral view D respective line drawing E anterior view F respective line drawing G associated scapulocoracoid in posterior view. Abbreviations: ch – choanae, cor – coracoid, d – dentary, dcr – dentary crest, m – maxillae, pl – palatine, pm – premaxillae, pmcr – premaxillaery crest, prid – palatal ridge, sca – scapula. Arrows indicate alveoli or teeth. Scale bar = 10 mm. Photos courtesy of The Natural History Museum.

crête écaille dessin musée +4
Obelignathus septimanicus holotype, MDE D30 from the 'Grès à Reptiles' Formation of Montouliers, southern France. Photographs and photogrammetric models of the right dentary in dorsal (A), anterior (B), medial (C), and lateral (D) views.
Taxons Obelignathus

Obelignathus septimanicus holotype, MDE D30 from the 'Grès à Reptiles' Formation of Montouliers, southern France. Photographs and photogrammetric models of the right dentary in dorsal (A), anterior (B), medial (C), and lateral (D) views.

France holotype Obelignathus formation
Barsboldia sicinskii gen. et sp.n., type specimen ZPAL MgD-11110,
Nemegt Formation, N Nemegt, Nemegt Basin, Gobi Desert, Mongolia
1. Right lateral view of sacrum, neural spine of last dorsal and two anterior caudals

included.
Taxons Barsboldia

Barsboldia sicinskii gen. et sp.n., type specimen ZPAL MgD-11110, Nemegt Formation, N Nemegt, Nemegt Basin, Gobi Desert, Mongolia 1. Right lateral view of sacrum, neural spine of last dorsal and two anterior caudals included.

Mongolie spécimen Barsboldia Nemegtia +1
Partial skull of Manidens condorensis from the Middle Jurassic Cañadón Asfalto Formation of Argentina. Skull reconstructions in lateral view. Dashed lines indicate estimated edges. Abbreviations: a angular antfo antorbital fossa asaf anterior surangular foramen be buccal emargination bo basioccipital bt basal tubera d dentary d1, 2, 11 dentary tooth 1, 2, 11 emfo external mandibular fossa f frontal gl glenoid gr groove j jugal jfl jugal flange jh jugal horn m maxilla m1, 11 maxillary tooth 1, 11 n nasal pd predentary pm premaxilla po postorbital pof postorbital fossa popr paroccipital process q quadrate qj quadratojugal ri ridge sa surangular sq squamosal.
Taxons Manidens

Partial skull of Manidens condorensis from the Middle Jurassic Cañadón Asfalto Formation of Argentina. Skull reconstructions in lateral view. Dashed lines indicate estimated edges. Abbreviations: a angular antfo antorbital fossa asaf anterior surangular foramen be buccal emargination bo basioccipital bt basal tubera d dentary d1, 2, 11 dentary tooth 1, 2, 11 emfo external mandibular fossa f frontal gl glenoid gr groove j jugal jfl jugal flange jh jugal horn m maxilla m1, 11 maxillary tooth 1, 11 n nasal pd predentary pm premaxilla po postorbital pof postorbital fossa popr paroccipital process q quadrate qj quadratojugal ri ridge sa surangular sq squamosal.

dent Argentine Jurassique Jurassique moyen +4
A 6.4 meter (21ft) Eurhinosaurus species specimen from the Holzmaden Formation, Baden-Wurttenburg, Germany.  ~185 million years old.
Taxons Eurhinosaurus

A 6.4 meter (21ft) Eurhinosaurus species specimen from the Holzmaden Formation, Baden-Wurttenburg, Germany. ~185 million years old.

Allemagne spécimen Eurhinosauria formation
Cast of a Scaphognathus crassirostris, a kind of pterosaur. On display as part of the exhibit "Pterosaurs: Flight in the Age of Dinosaurs" at the Cleveland Natural History Museum in Cleveland, Ohio, in the United States.
This animal lived about 150 million years ago. This fossil was found in the Solnhofen formation in Germany. This is a cast; the fossil itself is held by the Institute of Geology and Paleontology at the University of Bonn.
Taxons Scaphognathus

Cast of a Scaphognathus crassirostris, a kind of pterosaur. On display as part of the exhibit "Pterosaurs: Flight in the Age of Dinosaurs" at the Cleveland Natural History Museum in Cleveland, Ohio, in the United States. This animal lived about 150 million years ago. This fossil was found in the Solnhofen formation in Germany. This is a cast; the fossil itself is held by the Institute of Geology and Paleontology at the University of Bonn.

vol musée Allemagne États-Unis +7
Cimoliopterus cuvieri. Holotype NHMUK PV 39409 (Cenomanian / Turonian, Chalk Formation), anterior part of the rostrum A right lateral view B respective line drawing C ventral view D respective line drawing. Abbreviations: m – maxillae, pm – premaxillae, pmcr – premaxillary crest, prid – palatal ridge. Arrows and numbers indicate alveoli or teeth and their respective position. Scale bar = 10 mm. Photos courtesy of The Natural History Museum.
Taxons Cimoliopterus

Cimoliopterus cuvieri. Holotype NHMUK PV 39409 (Cenomanian / Turonian, Chalk Formation), anterior part of the rostrum A right lateral view B respective line drawing C ventral view D respective line drawing. Abbreviations: m – maxillae, pm – premaxillae, pmcr – premaxillary crest, prid – palatal ridge. Arrows and numbers indicate alveoli or teeth and their respective position. Scale bar = 10 mm. Photos courtesy of The Natural History Museum.

crête écaille dessin musée +5
Specimens of Galleonosaurus dorisae n. gen. n. sp. from the Flat Rocks Sandstone in the upper Barremian, Wonthaggi Formation, Gippsland Basin, southeastern Australia: (1–2) holotype (NMV P229196), left maxilla in lateral (1) and medial (2) views; (3) NMV P208178, left maxilla in lateral view; (4) NMV P212845, left maxilla in lateral view; (5) NMV P209977, left maxilla in lateral view; (6) NMV P186440, left maxilla in lateral view; (7) NMV 208113, right maxillary tooth in labial view. Scale bars = 10 mm (1–6); 1 mm (7).
Taxons Galleonosaurus

Specimens of Galleonosaurus dorisae n. gen. n. sp. from the Flat Rocks Sandstone in the upper Barremian, Wonthaggi Formation, Gippsland Basin, southeastern Australia: (1–2) holotype (NMV P229196), left maxilla in lateral (1) and medial (2) views; (3) NMV P208178, left maxilla in lateral view; (4) NMV P212845, left maxilla in lateral view; (5) NMV P209977, left maxilla in lateral view; (6) NMV P186440, left maxilla in lateral view; (7) NMV 208113, right maxillary tooth in labial view. Scale bars = 10 mm (1–6); 1 mm (7).

écaille dent Australie Barrémien +4
Azhdarcho sp., ZIN PH 56/43, distal fragment of a right ulna in proximal (a), ventral (b), posterior (c), dorsal (d), anterior (e), and distal (f, stereopair) views. This specimen is from the Tyulkili locality in the northeastern Aral Sea region of Kazakhstan; Zhirkindek Formation, Upper Cretaceous (upper Turonian – Coniacian). Abbreviations: das, dorsal articulation surface; ft, groove for flexor tendon; tub, tuberculum; vf, ventral fovea. Scale bar is 10 mm.
Taxons Azhdarcho

Azhdarcho sp., ZIN PH 56/43, distal fragment of a right ulna in proximal (a), ventral (b), posterior (c), dorsal (d), anterior (e), and distal (f, stereopair) views. This specimen is from the Tyulkili locality in the northeastern Aral Sea region of Kazakhstan; Zhirkindek Formation, Upper Cretaceous (upper Turonian – Coniacian). Abbreviations: das, dorsal articulation surface; ft, groove for flexor tendon; tub, tuberculum; vf, ventral fovea. Scale bar is 10 mm.

écaille Kazakhstan Coniacien Crétacé +4
Pelvis (fused ilia and pubes) of UALVP 48778, holotype of Hesperonychus elizabethae, from the late Campanian Dinosaur Park Formation of Alberta, Canada. Fossil collected by Dr. Elizabeth Nicholls. Prepared by Clive Coy. Digital photography by Nick Longrich. Additional digital editing by Nick Longrich.
Taxons Hesperonychus

Pelvis (fused ilia and pubes) of UALVP 48778, holotype of Hesperonychus elizabethae, from the late Campanian Dinosaur Park Formation of Alberta, Canada. Fossil collected by Dr. Elizabeth Nicholls. Prepared by Clive Coy. Digital photography by Nick Longrich. Additional digital editing by Nick Longrich.

bassin Canada Dinosaur Park Campanien +6
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Actualités

Épisode 173 : Forêt pétrifiée
États-Unis Chinle Trias supérieur Trias formation
Le parc national de la Forêt Pétrifiée, dans le nord-est de l'Arizona, aux États-Unis, est une plaque tournante de la paléontologie du Trias et présente des affleurements représentant 20 millions d'années de la formation Chinle du Trias supérieur. Les visiteurs s'émerveillent devant les arbres fossilisés colorés dont le parc tire son nom, mais toute une série d'animaux ont élu domicile dans ces forêts marécageuses il y a 225 millions d'années [&hellip
17/03/2026 palaeocast ⚙ Traduction automatique
Des fossiles de poissons vieux de 400 millions d'années révèlent comment la vie a commencé à s'installer sur terre
Australie Chine fossile formation crâne
Les scientifiques ont découvert de nouveaux indices sur certains des premiers poissons de la Terre, mettant ainsi en lumière les origines anciennes des vertébrés qui ont fini par s’installer sur terre. En réanalysant de mystérieux fossiles de la célèbre formation australienne Gogo et en étudiant un crâne de poisson-poumon récemment reconstruit, vieux de 410 millions d'années et provenant de Chine, les chercheurs révèlent comment ces créatures primitives ont évolué.
12/03/2026 sciencedaily ⚙ Traduction automatique
Une ancienne sécheresse aurait pu anéantir les vrais hobbits il y a 61 000 ans
extinction formation
Une sécheresse massive qui a duré des siècles pourrait avoir entraîné l’extinction des « hobbits » de Flores. Les enregistrements climatiques conservés dans les formations souterraines montrent que les précipitations ont chuté au moment même où la petite espèce humaine disparaissait. Dans le même temps, le nombre d’éléphants pygmées dont ils dépendaient a fortement diminué à mesure que les rivières s’asséchaient. Avec la disparition de la nourriture et de l'eau, les hobbits ont peut-être été repoussés et entrés dans leur dernier chapitre.
19/02/2026 sciencedaily-human-evo ⚙ Traduction automatique
Épisode 171 : Mosasaures d'eau douce
Hell Creek Mosasaurus formation
Nous plongeons dans les eaux douces de la formation Hell Creek. Oh cher
12/12/2025 palaeocast ⚙ Traduction automatique
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