formation

Bifurculapes laqueatus trackway (positive hyporelief, i.e. casts on the bottom surface of the bed) from the Early Jurassic East Berlin Formation (Hartford Basin) of Holyoke, Massachusetts (not the same specimen as this one). See coin (1 U.S. cent, 19.05 mm in diameter) for scale.

Bifurculapes laqueatus trackway (positive hyporelief, i.e. casts on the bottom surface of the bed) from the Early Jurassic East Berlin Formation (Hartford Basin) of Holyoke, Massachusetts (not the same specimen as this one). See coin (1 U.S. cent, 19.05 mm in diameter) for scale.

A trackway of the trace fossil Bifurculapes laqueatus (positive hyporelief, i.e. casts on the bottom surface of the bed) from the Early Jurassic East Berlin Formation (Hartford Basin) of Holyoke, Massachusetts (not the same specimen as this one). Scale is in cm.

Geological context of the Lower Cretaceous deposits of southeast England, focussing on the Purbeck Group and Wealden Supergroup. (A) Schematic geology of the Lower Cretaceous deposits of the Weald Sub-basin (southeast England), highlighting published spinosaurid finds (Charig & Milner, 1997; Salisbury & Naish, 2011; Turmine-Juhel et al., 2019). Based on Austen & Batten (2018: Fig. 2). Note that various additional spinosaurid teeth are known from the region but remain undescribed in detail (Fowler, 2007). (B) Simplified stratigraphic column of the Weald Group in southeast England, based on Batten & Austen (2011: Fig. 3.2). Note that the Grinstead Clay Formation, which subdivides the Tunbridge Wells Sands Formation in Batten & Austen (2011) and from which the “Suchosaurus cultridens” type specimen was discovered (Salisbury & Naish, 2011), is downgraded to a member of the latter formation in other works (Hopson, Wilkinson & Woods, 2008) and has not been included in this column. Spinosaurid silhouette courtesy of Dan Folkes (CC-BY 4.0).

Type section of the Cedar Mountain Formation, western side of Cedar Mountain, Emery County, Utah.

Morrison Formation (lower half), Jurassic-Cretacous boundary (K1 Unconformity) at red and orange paleosol, Cedar Mountain Formation (drab gray), and capping Naturita Formation. West of Dinosaur National Monument.

Brushy Basin Member of the Morrison Formation (multi-colored) unconformably overlain by the Yellow Cat Member of the Cedar Mountain Formation (drab color), Jessie's Twist, Emery County, Utah

Fossil eggs of the oospecies Macroelongatoolithus carlylei (believed to be the eggs of giant oviraptorosaurs) from the Cedar Mountain Formation of North America. At the SECU Dinolab of the North Carolina Museum of Natural Sciences

View east towards the Quarry Visitor Center at Dinosaur National Monument. The boundary between the Morrison Formation (Brushy Basin Member) and the Cedar Mountain Formation is just above the highest red bed.

Typical exposure of the Cedar Mountain Formation overlying the Morrison Formation, south of Green River Utah

This image shows the Cedar Mountain Formation near its type section at Buckhorn Reservoir, Utah. The formation is capped by a very thin bed of Dakota Formation. The Cedar Mountain Formation is a Cretaceous fluvial formation, about 98 million years old, that was deposited in the foreland basin of the Sevier Mountains just before it was flooded by the ocean to form the Western Interior Seaway.

Diagram depicting the currently named Dinosauria from the Early Cretaceous Cedar Mountain Formation of Utah. Dinosaur taxa: Cedarosaurus weiskopfae → Grand County locality.[1] Length = 15 meters.[2] Cedrorestes crichtoni → Dave's Camp Site locality.[3] Length = 6 meters.[4] Falcarius utahensis → Crystal Geyser Quarry & Suarez Site/Quarry localities.[5] Length = 5 meters.[5] Gastonia burgei → Gaston Quarry, Dalton Wells, Doelling's Bowl localities.[6] Length = 5 meters.[2] Geminiraptor suarezarum → Suarez Site/Quarry locality.[7] Length = 1.5 meters.[4] Hippodraco scutodens → Andrew's Site locality.[8] Length = 4.5 meters.[8] Iguanacolossus fortis → Don's Ridge & Doelling's Bowl localities.[8][6] Length = 9 meters.[8] Martharaptor greenriverensis → Hayden-Corbett Site.[9] Length = Extrapolated after relatives. Mierasaurus bobyoungi → Doelling’s Bowl locality.[10] Length = ∼9 meters.[10][11] Moabosaurus utahensis → Dalton Wells locality.[12] Length = 9.75 meters.[12] Nedcolbertia justinhofmanni → Dalton Wells & Gaston Quarry localities.[13] Length = ∼3 meters.[13] Utahraptor ostrommaysi → Dalton Wells, Gaston Quarry, Stikes Quarry, Utahraptor Ridge localities.[14][6] Length = 5.5 meters.[2] Yurgovuchia doellingi → Don’s Place - Doelling’s Bowl locality.[15] Length = 2.5 meters.[15] References ↑ (1999). "New sauropod from the Lower Cretaceous of Utah, USA". Oryctos 2: 21–37. ↑ a b c (2016) The Princeton Field Guide to Dinosaurs (2nd ed.), Princeton: Princeton University Press ISBN: 9780691167664. ↑ "A possible new basal hadrosaur from the Lower Cretaceous Cedar Mountain Formation of Eastern Utah" in (2007) Horns and Beaks: Ceratopsian and Ornithopod Dinosaurs, Bloomington and Indianapolis: Indiana University Press, pp. 79–89 DOI: 10.2307/j.ctt1zxz1md.10. ISBN: 0-253-34817-X. ↑ a b (2007) Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages, Random House ISBN: 9780375824197. Genus List for Holtz 2012 Weight Information ↑ a b (2010). "Osteology of Falcarius utahensis (Dinosauria: Theropoda): characterizing the anatomy of basal therizinosaurs". Zoological Journal of the Linnean Society 158 (1): 196–230. DOI:10.1111/j.1096-3642.2009.00464.x. ↑ a b c (2016). "The Lower Cretaceous in East-Central Utah—The Cedar Mountain Formation and its Bounding Strata". Geology of the Intermoutain West 3: 1-130. ↑ (2010). "A New Troodontid Theropod Dinosaur from the Lower Cretaceous of Utah". PLOS ONE 5 (12): e14329. DOI:10.1371/journal.pone.0014329. PMID 21179513. PMC: 3002269. ↑ a b c d (2010). "New Basal Iguanodonts from the Cedar Mountain Formation of Utah and the Evolution of Thumb-Spiked Dinosaurs". PLOS ONE 5 (11): e14075. DOI:10.1371/journal.pone.0014075. PMID 21124919. PMC: 2989904. ↑ (2012). "Martharaptor greenriverensis, a New Theropod Dinosaur from the Lower Cretaceous of Utah". PLOS ONE 7 (8): e43911. DOI:10.1371/journal.pone.0043911. PMID 22952806. PMC: 3430620. ↑ a b (2017). "Descendants of the Jurassic turiasaurs from Iberia found refuge in the Early Cretaceous of western USA". Scientific Reports 7 (1): 14311. DOI:10.1038/s41598-017-14677-2. PMID 29085006. PMC: 5662694. ↑ (2020) Dinosaur Facts and Figures: The Sauropods and Other Sauropodomorphs, Princeton: Princeton University Press ISBN: 9780691202976. ↑ a b (2017). "Moabosaurus Utahensis, N. Gen., N. SP., A New Sauropod From The Early Cretaceous (Aptian) of North America". Contributions from the Museum of Paleontology, University of Michigan 32 (11): 189–243. ↑ a b (1998). "A small coelurosaurian theropod from the Yellow Cat Member of the Cedar Mountain Formation (Lower Cretaceous, Barremian) of eastern Utah". New Mexico Museum of Natural History and Science Bulletin 14: 239–248. ↑ (1993). "A large dromaeosaurid (Theropoda) from the Lower Cretaceous of Eastern Utah". Hunteria 2 (10): 1–16. ↑ a b (2012). "New Dromaeosaurids (Dinosauria: Theropoda) from the Lower Cretaceous of Utah, and the Evolution of the Dromaeosaurid Tail". PLOS ONE 7 (5): e36790. DOI:10.1371/journal.pone.0036790. PMID 22615813. PMC: 3352940.

Holotype of Khunnuchelys lophorhothon sp. nov. from Baybishe, northeastern Aral Sea region, Kazakhstan, Bostobe Formation, Santonian–early Campanian, Late Cretaceous, ZIN PH 5/55, a partial skull; in dorsal (A), ventral (B), lateral (C), and anterior (D) views. Photographs (A1–D1), explanatory drawings (A2–D2).

World map showing Late Cretaceous metatherian locales. Europe 1. Font-de-Benon quarry, Archingeay-Les Nouillers (Cenomanian, Late Cretaceous), Charente-Maritime, southwestern France (Vullo et al. 2009) 2. Valkenburg Member, Maastricht Formation (late Maastrichtian, Late Cretaceous), southern Limburg, The Netherlands (Martin et al. 2005) Asia 3. Yixian Formation, China (Barremian, Early Cretaceous) 4. Bissekty Formation, Kyzylkum Desert, Uzbekistan (Turonian, Late Cretaceous) 5. Darbasa Formation, southern Kazakhstan (Campanian, Late Cretaceous) 6. Grey Mesa locality (Averianov 1997) 7. Barun Goyot Formation, Umuni Gobi, Mongolia (Campanian, Late Cretaceous) 8. Nemegt Formation, Omnogov, Mongolia (Maastrichtian, Late Cretaceous) 9. Djadokhta Formation, Mongolia (Campanian, Late Cretaceous) North America Alaska 9. Prince Creek Formation, Alaska (early Maastrichtian, Late Cretaceous) Alberta and Saskatchewan, Canada 10. Milk River Formation, southern Alberta, Canada (late Santonian, Late Cretaceous) 11. Oldman Formation, southern Alberta, Canada (Campanian, Late Cretaceous) 12. Dinosaur Park Formation, southern Alberta, Canada (late Campanian, Late Cretaceous) 13. Horseshoe Canyon Formation, southern Alberta, Canada (early Maastrichtian, Late Cretaceous) 14. St. Mary River Formation, Alberta and northwestern Montana (early Maastrichtian, Late Cretaceous) 15. Scollard Formation, Alberta (late Maastrichtian, Late Cretaceous) 16. Frenchman Formation, Saskatchewan (late Maastrichtian, Late Cretaceous) Montana and North Dakota 17. Judith River Formation (late Campanian, Late Cretaceous) 18. Two Medicine Formation (late Campanian, Late Cretaceous) 19. Hell Creek Formation, Montana and North Dakota (late Maastrichtian, Late Cretaceous) South Dakota 20. Fox Hills Formation, South Dakota (late Maastrichtian, Late Cretaceous) 21. Hell Creek Formation, South Dakota (late Maastrichtian, Late Cretaceous) Wyoming 22. “Mesa Verde Formation” (late Campanian, Late Cretaceous) 23. Lance Formation, Wyoming (late Maastrichtian, Late Cretaceous) 24. Ferris Formation, Wyoming (late Maastrichtian, Late Cretaceous) Utah 25. Cedar Mountain Formation (Albian-Cenomanian) 26. Dakota Formation fauna (late Cenomanian, Late Cretaceous) 27. Smoky Hollow Member, Straight Cliffs Formation (Turonian, Late Cretaceous) 28. John Henry Member, Straight Cliffs Formation (Coniacian-Santonian, Late Cretaceous) 29. Wahweap Formation (early-middle Campanian, Late Cretaceous) 30. Kaiparowits Formation (late Campanian, Late Cretaceous) 31. Iron Springs Formation fauna, southern Utah (Turonian – Santonian, Late Cretaceous) 32. North Horn Formation, Utah (late Maastrichtian, Late Cretaceous) Colorado 33. Williams Fork Formation, Colorado (late Campanian-early Maastrichtian) 34. Laramie Formation, northeastern Colorado (late Maastrichtian, Late Cretaceous) Baja California Del Norte, Mexico 35. El Gallo Formation (late Campanian, Late Cretaceous) (Clemens 1980; Lillegraven 1972; Lillegraven 1976) New Mexico 36. Fruitland and lower Kirtland Formation, San Juan Basin (late Campanian, Late Cretaceous) 37. Naashoibito Member, Kirtland Formation, New Mexico (late Maastrichtian, Late Cretaceous) Oklahoma 38. Antlers Formation, Texas and Oklahoma (Aptian-Albian, Early Cretaceous) Texas 39. Aguja Formation, West Texas (late Campanian, Late Cretaceous) New Jersey 40. Marshalltown Formation, New Jersey (Campanian, Late Cretaceous) Szalay 1994)

World map showing Late Cretaceous metatherian locales. Europe 1. Font-de-Benon quarry, Archingeay-Les Nouillers (Cenomanian, Late Cretaceous), Charente-Maritime, southwestern France (Vullo et al. 2009) 2. Valkenburg Member, Maastricht Formation (late Maastrichtian, Late Cretaceous), southern Limburg, The Netherlands (Martin et al. 2005) Asia 3. Yixian Formation, China (Barremian, Early Cretaceous) 4. Bissekty Formation, Kyzylkum Desert, Uzbekistan (Turonian, Late Cretaceous) 5. Darbasa Formation, southern Kazakhstan (Campanian, Late Cretaceous) 6. Grey Mesa locality (Averianov 1997) 7. Barun Goyot Formation, Umuni Gobi, Mongolia (Campanian, Late Cretaceous) 8. Nemegt Formation, Omnogov, Mongolia (Maastrichtian, Late Cretaceous) 9. Djadokhta Formation, Mongolia (Campanian, Late Cretaceous) North America Alaska 9. Prince Creek Formation, Alaska (early Maastrichtian, Late Cretaceous) Alberta and Saskatchewan, Canada 10. Milk River Formation, southern Alberta, Canada (late Santonian, Late Cretaceous) 11. Oldman Formation, southern Alberta, Canada (Campanian, Late Cretaceous) 12. Dinosaur Park Formation, southern Alberta, Canada (late Campanian, Late Cretaceous) 13. Horseshoe Canyon Formation, southern Alberta, Canada (early Maastrichtian, Late Cretaceous) 14. St. Mary River Formation, Alberta and northwestern Montana (early Maastrichtian, Late Cretaceous) 15. Scollard Formation, Alberta (late Maastrichtian, Late Cretaceous) 16. Frenchman Formation, Saskatchewan (late Maastrichtian, Late Cretaceous) Montana and North Dakota 17. Judith River Formation (late Campanian, Late Cretaceous) 18. Two Medicine Formation (late Campanian, Late Cretaceous) 19. Hell Creek Formation, Montana and North Dakota (late Maastrichtian, Late Cretaceous) South Dakota 20. Fox Hills Formation, South Dakota (late Maastrichtian, Late Cretaceous) 21. Hell Creek Formation, South Dakota (late Maastrichtian, Late Cretaceous) Wyoming 22. “Mesa Verde Formation” (late Campanian, Late Cretaceous) 23. Lance Formation, Wyoming (late Maastrichtian, Late Cretaceous) 24. Ferris Formation, Wyoming (late Maastrichtian, Late Cretaceous) Utah 25. Cedar Mountain Formation (Albian-Cenomanian) 26. Dakota Formation fauna (late Cenomanian, Late Cretaceous) 27. Smoky Hollow Member, Straight Cliffs Formation (Turonian, Late Cretaceous) 28. John Henry Member, Straight Cliffs Formation (Coniacian-Santonian, Late Cretaceous) 29. Wahweap Formation (early-middle Campanian, Late Cretaceous) 30. Kaiparowits Formation (late Campanian, Late Cretaceous) 31. Iron Springs Formation fauna, southern Utah (Turonian – Santonian, Late Cretaceous) 32. North Horn Formation, Utah (late Maastrichtian, Late Cretaceous) Colorado 33. Williams Fork Formation, Colorado (late Campanian-early Maastrichtian) 34. Laramie Formation, northeastern Colorado (late Maastrichtian, Late Cretaceous) Baja California Del Norte, Mexico 35. El Gallo Formation (late Campanian, Late Cretaceous) (Clemens 1980; Lillegraven 1972; Lillegraven 1976) New Mexico 36. Fruitland and lower Kirtland Formation, San Juan Basin (late Campanian, Late Cretaceous) 37. Naashoibito Member, Kirtland Formation, New Mexico (late Maastrichtian, Late Cretaceous) Oklahoma 38. Antlers Formation, Texas and Oklahoma (Aptian-Albian, Early Cretaceous) Texas 39. Aguja Formation, West Texas (late Campanian, Late Cretaceous) New Jersey 40. Marshalltown Formation, New Jersey (Campanian, Late Cretaceous) Szalay 1994)

Paleogeography and paleoclimate of the Late Jurassic - 150 Ma with dinosaur fossil localities: A = Tendaguru Formation, Tanzania C1 = Shishugou & Kalazha Formations, China C2 = Shangshaximiao (Upper Shaximiao) Formation, China E1 = Sables de Glos, Argiles d’Octeville, Marnes de Bléville, Kimmeridge Clay, Calcareous Grit, Corallian Oolite, Oxford Clay, Portland Stone, England & France E2 = Villar del Arzobispo, Alcobaça, Guimarota, Sobral, Amoreira-Porto Novo, Bombarral, Freixial, Lourinhã Formations, Spain & Portugal M1-6 = Morrison Formation, United States S1 = Toquí & Cañadón Calcáreo Formations, Chile & Argentina