holotype

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The holotype and only known specimen of the hauffiopterygian leptonectid, Xiphodracon goldencapensis (ROM VP52596) from Golden Cap, between Charmouth and Seatown, Dorset, UK. The skeleton is exposed in ventrolateral view. The skull has been fully prepared free of matrix whereas most of the skeleton is still in matrix. The left (upper) forefin has been prepared so that it is three-dimensionally preserved and projects upwards. Scale bar represents 20 cm.
Taxons Xiphodracon

The holotype and only known specimen of the hauffiopterygian leptonectid, Xiphodracon goldencapensis (ROM VP52596) from Golden Cap, between Charmouth and Seatown, Dorset, UK. The skeleton is exposed in ventrolateral view. The skull has been fully prepared free of matrix whereas most of the skeleton is still in matrix. The left (upper) forefin has been prepared so that it is three-dimensionally preserved and projects upwards. Scale bar represents 20 cm.

écaille Royaume-Uni holotype spécimen +5
Skeletal reconstruction of Siamraptor suwati. Cranial elements were scaled to fit in with the holotype (surangular). Human size = 1.8 m. Scale bar = 1 m. Human silhouette has been added to the original image.
Taxons Siamraptor

Skeletal reconstruction of Siamraptor suwati. Cranial elements were scaled to fit in with the holotype (surangular). Human size = 1.8 m. Scale bar = 1 m. Human silhouette has been added to the original image.

écaille holotype Siamraptor
Skeletal reconstruction of Ahshislepelta minor, a small ankylosaur from the Late Cretaceous Kirtland Formation of New Mexico. While originally recovered as an ankylosaurid related to Gastonia, later analyses favored a nodosaurid position which this is based on. Based on the holotype SMP VP-1930, consisting of fragmentary shoulder, forelimb, and vertebral elements as well as several osteoderms. Unknown material filled in using Niobrarasaurus, Pawpawsaurus, Silvisaurus, Sauropelta, and Borealopelta. Total length is approximately 4.9 m through the centra.
Notes: Osteoderm placement somewhat speculative, not all osteoderms and vertebral fragments are figured.

References: Burns & Sullivan, 2011.
Taxons Ahshislepelta

Skeletal reconstruction of Ahshislepelta minor, a small ankylosaur from the Late Cretaceous Kirtland Formation of New Mexico. While originally recovered as an ankylosaurid related to Gastonia, later analyses favored a nodosaurid position which this is based on. Based on the holotype SMP VP-1930, consisting of fragmentary shoulder, forelimb, and vertebral elements as well as several osteoderms. Unknown material filled in using Niobrarasaurus, Pawpawsaurus, Silvisaurus, Sauropelta, and Borealopelta. Total length is approximately 4.9 m through the centra. Notes: Osteoderm placement somewhat speculative, not all osteoderms and vertebral fragments are figured. References: Burns & Sullivan, 2011.

Mexique Kirtland Crétacé Crétacé supérieur +13
Aetodactylus halli holotype.
Taxons Aetodactylus

Aetodactylus halli holotype.

holotype Aetodactylus
Skeletal composite of the chilean titanosaur Atacamatitan chilensis. A fragmented sauropod known from the holotype SGO-PV-961, found in the Atacama Desert. The preserved elements consist of:[1]
Right femur, the proximal end of a humerus, two dorsal vertebrae, posterior caudal vertebrae, dorsal ribs and a possibly fragmentary element of the sternum, other fragmented caudal vertebrae and indeterminate bones.[1]
Due to the fragmented nature of the holotype, most cladistic analyzes exclude Atacamatitan. Nevertheless, in 2012, Rubilar-Rogers and Gutstein conducted a preliminary cladistic analysis which placed Atacamatitan within the Lithostrotia.[2]
The original description lacks a lateral view for the humerus and femur, the lateral view for the humerus was based on Alamosaurus and Opisthocoelicaudia, since it shows resemblance with them. The lateral view of the femur is based on Mendozasaurus for the same reason.

Color Key
  Known
  Unknown
Taxons Mendozasaurus

Skeletal composite of the chilean titanosaur Atacamatitan chilensis. A fragmented sauropod known from the holotype SGO-PV-961, found in the Atacama Desert. The preserved elements consist of:[1] Right femur, the proximal end of a humerus, two dorsal vertebrae, posterior caudal vertebrae, dorsal ribs and a possibly fragmentary element of the sternum, other fragmented caudal vertebrae and indeterminate bones.[1] Due to the fragmented nature of the holotype, most cladistic analyzes exclude Atacamatitan. Nevertheless, in 2012, Rubilar-Rogers and Gutstein conducted a preliminary cladistic analysis which placed Atacamatitan within the Lithostrotia.[2] The original description lacks a lateral view for the humerus and femur, the lateral view for the humerus was based on Alamosaurus and Opisthocoelicaudia, since it shows resemblance with them. The lateral view of the femur is based on Mendozasaurus for the same reason. Color Key   Known   Unknown

os humérus description holotype +6
Dinosaur National Monument is a United States National Monument located on the southeast flank of the Uinta Mountains on the border between Colorado and Utah at the confluence of the Green and Yampa Rivers. Although most of the monument area is in Moffat County, Colorado, the Dinosaur Quarry is located in Utah just to the north of the town of Jensen, Utah.
The nearest communities are Jensen, Utah, and Dinosaur, Colorado. The park contains over 800 paleontological sites and has fossils of dinosaurs including Allosaurus, Deinonychus, Abydosaurus (a nearly complete skull, lower jaws and first four neck vertebrae of the specimen DINO 16488 found here at the base of the Mussentuchit Member of the Cedar Mountain Formation is the holotype for the description) and various long-neck, long-tail sauropods. It was declared a National Monument on October 4, 1915.
The rock layer enclosing the fossils is a sandstone and conglomerate bed of alluvial or river bed origin known as the Morrison Formation from the Jurassic Period some 150 million years old. The dinosaurs and other ancient animals were carried by the river system which eventually entombed their remains in Utah.
The pile of sediments were later buried and lithified into solid rock. The layers of rock were later uplifted and tilted to their present angle by the mountain building forces that formed the Uintas during the Laramide orogeny. The relentless forces of erosion exposed the layers at the surface to be found by paleontologists.
The dinosaur fossil beds (bone beds) were discovered in 1909 by Earl Douglass, a paleontologist working and collecting for the Carnegie Museum of Natural History. He and his crews excavated thousands of fossils and shipped them back to the museum in Pittsburgh, Pennsylvania for study and display. President Woodrow Wilson proclaimed the dinosaur beds as Dinosaur National Monument in 1915. The monument boundaries were expanded in 1938 from the original 80-acre (320,000 m2) tract surrounding the dinosaur quarry in Utah, to its present extent of over 200,000 acres (800 km²) in Utah and Colorado, encompassing the spectacular river canyons of the Green and Yampa.
Though lesser-known than the fossil beds, the petroglyphs in Dinosaur National Monument are another treasure the monument holds. Due to problems with vandals, many of the sites are not listed on area maps.
The "Wall of Bones" located within the Dinosaur Quarry building in the park consists of a steeply tilted (67° from horizontal) rock layer which contains hundreds of dinosaur fossils. The enclosing rock has been chipped away to reveal the fossil bones intact for public viewing. In July 2006, the Quarry Visitor Center was closed due to structural problems that since 1957 had plagued the building because it was built on unstable clay. The decision was made to build a new facility elsewhere in the monument to house the visitor center and administrative functions, making it easier to resolve the structural problems of the quarry building while still retaining a portion of the historic Mission 66 era exhibit hall. It was announced in April 2009 that Dinosaur National Monument would receive $13.1 million to refurbish and reopen the gallery as part of the Obama administration's $750 billion stimulus plan. The Park Service successfully rebuilt the Quarry Exhibit Hall, supporting its weight on 70-foot steel micropile columns that extend to the bedrock below the unstable clay. The Dinosaur Quarry was reopened in Fall 2011.
en.wikipedia.org/wiki/Dinosaur_National_Monument

en.wikipedia.org/wiki/Wikipedia:Text_of_Creative_Commons_...
Taxons Abydosaurus

Dinosaur National Monument is a United States National Monument located on the southeast flank of the Uinta Mountains on the border between Colorado and Utah at the confluence of the Green and Yampa Rivers. Although most of the monument area is in Moffat County, Colorado, the Dinosaur Quarry is located in Utah just to the north of the town of Jensen, Utah. The nearest communities are Jensen, Utah, and Dinosaur, Colorado. The park contains over 800 paleontological sites and has fossils of dinosaurs including Allosaurus, Deinonychus, Abydosaurus (a nearly complete skull, lower jaws and first four neck vertebrae of the specimen DINO 16488 found here at the base of the Mussentuchit Member of the Cedar Mountain Formation is the holotype for the description) and various long-neck, long-tail sauropods. It was declared a National Monument on October 4, 1915. The rock layer enclosing the fossils is a sandstone and conglomerate bed of alluvial or river bed origin known as the Morrison Formation from the Jurassic Period some 150 million years old. The dinosaurs and other ancient animals were carried by the river system which eventually entombed their remains in Utah. The pile of sediments were later buried and lithified into solid rock. The layers of rock were later uplifted and tilted to their present angle by the mountain building forces that formed the Uintas during the Laramide orogeny. The relentless forces of erosion exposed the layers at the surface to be found by paleontologists. The dinosaur fossil beds (bone beds) were discovered in 1909 by Earl Douglass, a paleontologist working and collecting for the Carnegie Museum of Natural History. He and his crews excavated thousands of fossils and shipped them back to the museum in Pittsburgh, Pennsylvania for study and display. President Woodrow Wilson proclaimed the dinosaur beds as Dinosaur National Monument in 1915. The monument boundaries were expanded in 1938 from the original 80-acre (320,000 m2) tract surrounding the dinosaur quarry in Utah, to its present extent of over 200,000 acres (800 km²) in Utah and Colorado, encompassing the spectacular river canyons of the Green and Yampa. Though lesser-known than the fossil beds, the petroglyphs in Dinosaur National Monument are another treasure the monument holds. Due to problems with vandals, many of the sites are not listed on area maps. The "Wall of Bones" located within the Dinosaur Quarry building in the park consists of a steeply tilted (67° from horizontal) rock layer which contains hundreds of dinosaur fossils. The enclosing rock has been chipped away to reveal the fossil bones intact for public viewing. In July 2006, the Quarry Visitor Center was closed due to structural problems that since 1957 had plagued the building because it was built on unstable clay. The decision was made to build a new facility elsewhere in the monument to house the visitor center and administrative functions, making it easier to resolve the structural problems of the quarry building while still retaining a portion of the historic Mission 66 era exhibit hall. It was announced in April 2009 that Dinosaur National Monument would receive $13.1 million to refurbish and reopen the gallery as part of the Obama administration's $750 billion stimulus plan. The Park Service successfully rebuilt the Quarry Exhibit Hall, supporting its weight on 70-foot steel micropile columns that extend to the bedrock below the unstable clay. The Dinosaur Quarry was reopened in Fall 2011. en.wikipedia.org/wiki/Dinosaur_National_Monument en.wikipedia.org/wiki/Wikipedia:Text_of_Creative_Commons_...

os description musée États-Unis +13
Life restoration of the Triassic ichthyosaur Callawayia neoscapularis. Three specimens of this ichthyosaur are known, the holotype, ROM 41993, and two referred specimens, TMP 94.380.11 and 94.382.2. The skull is primarily based on ROM 41993, cross-checked against TMP 94.380.11 and TMP 94.382.2. The vertebral column is based primarily on TMP 94.382.2 as it is the most complete of these specimens, while the ribs were based on ROM 41993. The forelimbs were mainly based on those of ROM 41993, with TMP 94.380.11 used to determine their breadth. The hindlimbs were based on TMP 94.380.11, especially the more complete right hindlimb.
ROM 41993 was cross-scaled with TMP 94.380.11 by the dimensions of the forelimb epipodials, which produced similar vertebral dimensions. The two TMP specimens were cross-scaled based on femoral length, also producing similar vertebral dimensions. Nicholls & Manabe (2001) stated that no wedge-shaped caudal centra supporting a tailbend were found and that there was no evidence of a bend being present, though considered that they might have existed in the gap in the preserved caudals. Since various other Triassic ichthyosaurs have since been found to have tail bends, one was illustrated here. A modest downturn of roughly 15° was illustrated, comparable to that in Guanlingsaurus, and the location of the bend within the gap in the preserved vertebrae matches well with the location of the bend in Guizhouichthyosaurus.

References
McGowan, C. (1994). "A new species of Shastasaurus (Reptilia: Ichthyosauria) from the Triassic of British Columbia: The most complete exemplar of the genus". Journal of Vertebrate Paleontology 14 (2): 168–179. DOI:10.1080/02724634.1994.10011550.
Nicholls, E. L.; Manabe, M. (2001). "A new genus of ichthyosaur from the Late Triassic Pardonet Formation of British Columbia: Bridging the Triassic-Jurassic gap". Canadian Journal of Earth Sciences 38 (6): 983–1002.
Ji, C.; Jiang, D.Y.; Hao, W.; Sun, Y. (2011). "True tailbend occurred in the Late Triassic: Evidence from ichthyosaur skeletons of South China". Acta Scientiarum Naturalium Universitatis Pekinensis 47 (2): 309–314.
Shang, Q. H.; Li, C. (2009). "On the occurrence of the ichthyosaur Shastasaurus in the Guanling biota (Late Triassic), Guizhou, China". Vertebrata PalAsiatica 47 (3): 178–193.
Taxons Guanlingsaurus

Life restoration of the Triassic ichthyosaur Callawayia neoscapularis. Three specimens of this ichthyosaur are known, the holotype, ROM 41993, and two referred specimens, TMP 94.380.11 and 94.382.2. The skull is primarily based on ROM 41993, cross-checked against TMP 94.380.11 and TMP 94.382.2. The vertebral column is based primarily on TMP 94.382.2 as it is the most complete of these specimens, while the ribs were based on ROM 41993. The forelimbs were mainly based on those of ROM 41993, with TMP 94.380.11 used to determine their breadth. The hindlimbs were based on TMP 94.380.11, especially the more complete right hindlimb. ROM 41993 was cross-scaled with TMP 94.380.11 by the dimensions of the forelimb epipodials, which produced similar vertebral dimensions. The two TMP specimens were cross-scaled based on femoral length, also producing similar vertebral dimensions. Nicholls & Manabe (2001) stated that no wedge-shaped caudal centra supporting a tailbend were found and that there was no evidence of a bend being present, though considered that they might have existed in the gap in the preserved caudals. Since various other Triassic ichthyosaurs have since been found to have tail bends, one was illustrated here. A modest downturn of roughly 15° was illustrated, comparable to that in Guanlingsaurus, and the location of the bend within the gap in the preserved vertebrae matches well with the location of the bend in Guizhouichthyosaurus. References McGowan, C. (1994). "A new species of Shastasaurus (Reptilia: Ichthyosauria) from the Triassic of British Columbia: The most complete exemplar of the genus". Journal of Vertebrate Paleontology 14 (2): 168–179. DOI:10.1080/02724634.1994.10011550. Nicholls, E. L.; Manabe, M. (2001). "A new genus of ichthyosaur from the Late Triassic Pardonet Formation of British Columbia: Bridging the Triassic-Jurassic gap". Canadian Journal of Earth Sciences 38 (6): 983–1002. Ji, C.; Jiang, D.Y.; Hao, W.; Sun, Y. (2011). "True tailbend occurred in the Late Triassic: Evidence from ichthyosaur skeletons of South China". Acta Scientiarum Naturalium Universitatis Pekinensis 47 (2): 309–314. Shang, Q. H.; Li, C. (2009). "On the occurrence of the ichthyosaur Shastasaurus in the Guanling biota (Late Triassic), Guizhou, China". Vertebrata PalAsiatica 47 (3): 178–193.

Chine Jurassique Trias supérieur Trias +12
Fostoria dhimbangunmal (3D digital rendering of LRF 3050; holotype) braincase in I, dorsal and J, ventral views. Scale bar = 10 cm.
Taxons Fostoria

Fostoria dhimbangunmal (3D digital rendering of LRF 3050; holotype) braincase in I, dorsal and J, ventral views. Scale bar = 10 cm.

écaille holotype Fostoria
Shetwemys, Shell remains of the podocnemidid turtle S. fajumensis (Erymnochelyini). (a–c) SMNS 11233/2, partial carapace, in dorsal (a), ventral (b), and left lateral (c) views. (d) Ventral view of the anterior lobe the holotype of the species, currently lost, based on the fig. 2C in plate 8 of Andrews (1903). (e–g) SMNS 12647, plastron, in ventral (e), dorsal (f), and left lateral (g) views. (g’) corresponds to an enlarged photograph of the posterior plastral lobe, in left lateral view, in which the thickness in the regions close to the hypo-xiphiplastral suture (in blue), between the pelvic scars (in green), and at the level of the anal notch (in red), have been represented by arrows (h–i), SMNS 12646, plastron, in ventral (h) and dorsal (i) views. Gebel Quatrani Formation, Fayum depression, Egypt, Lower Oligocene (Rupelian)

Shetwemys, Shell remains of the podocnemidid turtle S. fajumensis (Erymnochelyini). (a–c) SMNS 11233/2, partial carapace, in dorsal (a), ventral (b), and left lateral (c) views. (d) Ventral view of the anterior lobe the holotype of the species, currently lost, based on the fig. 2C in plate 8 of Andrews (1903). (e–g) SMNS 12647, plastron, in ventral (e), dorsal (f), and left lateral (g) views. (g’) corresponds to an enlarged photograph of the posterior plastral lobe, in left lateral view, in which the thickness in the regions close to the hypo-xiphiplastral suture (in blue), between the pelvic scars (in green), and at the level of the anal notch (in red), have been represented by arrows (h–i), SMNS 12646, plastron, in ventral (h) and dorsal (i) views. Gebel Quatrani Formation, Fayum depression, Egypt, Lower Oligocene (Rupelian)

Égypte Oligocène Rupélien holotype +2
Close up of the Eulithomyrmex rugosus holotype head.  Museum of Comparative Zoology  specimen UCM17019.
Priabonian; Florissant Formation, Colorado, USA
Intervalles Priabonian

Close up of the Eulithomyrmex rugosus holotype head. Museum of Comparative Zoology specimen UCM17019. Priabonian; Florissant Formation, Colorado, USA

musée États-Unis Priabonien holotype +2
The Holotype batomorph tooth of Antiquaobatis grimmenensis from the Late Pliensbachian (spinatum) of Grimmen (Mecklenburg-Western Pomerania, Germany).
Intervalles Pliensbachian

The Holotype batomorph tooth of Antiquaobatis grimmenensis from the Late Pliensbachian (spinatum) of Grimmen (Mecklenburg-Western Pomerania, Germany).

dent Allemagne Pliensbachien holotype
New species of Mesozoic benthic foraminifera from the former British Petroleum micropalaeontology collection, Ophthalmidium dracomaris n. sp.
(1) holotype (NHMUK PM PF 74496); (2–5) paratypes (NHMUK PM PF 74497–74500). (8–9) Eobigenerina calloviensis n. sp.: (8) holotype (NHMUK PM PF 74504); (9) paratype (NHMUK PM PF 74505). (10–12) Trochammina fordonensis n. sp.: (10) holotype (NHMUK PM PF 74501); (11–12) paratype (NHMUK PM PF 74502–74503). (13–15) Arenoturrispirillina swiecickii n. sp.: (13) holotype (NHMUK PM PF 74506); (14–15) paratype (NHMUK PM PF 74507 + NHMUK PM PF 74552). (16–17) Ataxophragmium mariae n. sp.: (16) holotype (NHMUK PM PF 74553); (17) paratype (NHMUK PM PF 74554).

New species of Mesozoic benthic foraminifera from the former British Petroleum micropalaeontology collection, Ophthalmidium dracomaris n. sp. (1) holotype (NHMUK PM PF 74496); (2–5) paratypes (NHMUK PM PF 74497–74500). (8–9) Eobigenerina calloviensis n. sp.: (8) holotype (NHMUK PM PF 74504); (9) paratype (NHMUK PM PF 74505). (10–12) Trochammina fordonensis n. sp.: (10) holotype (NHMUK PM PF 74501); (11–12) paratype (NHMUK PM PF 74502–74503). (13–15) Arenoturrispirillina swiecickii n. sp.: (13) holotype (NHMUK PM PF 74506); (14–15) paratype (NHMUK PM PF 74507 + NHMUK PM PF 74552). (16–17) Ataxophragmium mariae n. sp.: (16) holotype (NHMUK PM PF 74553); (17) paratype (NHMUK PM PF 74554).

Mésozoïque holotype nouvelle espèce
Eekaulostomus cuevasae gen. and sp. nov. from the Danian marine deposits of the Belisario Domínguez quarry, near Palenque, Chiapas, southeastern Mexico. 1, General view of IGM 4716, holotype and single specimen known. 2, Idealized line drawing of the same specimen.

Eekaulostomus cuevasae gen. and sp. nov. from the Danian marine deposits of the Belisario Domínguez quarry, near Palenque, Chiapas, southeastern Mexico. 1, General view of IGM 4716, holotype and single specimen known. 2, Idealized line drawing of the same specimen.

dessin Mexique Danien holotype +1
Palaeohypsodontus zinensis sp. nov., late Oligocene (Chattian), Lundo J2, Bugti Hills, Balochistan, Pakistan. Holotype (ISEM DBJ2−A1). A. Stereophoto of occlusal view. B. Stereophoto of labial view. Scale bars 1 cm.

Palaeohypsodontus zinensis sp. nov., late Oligocene (Chattian), Lundo J2, Bugti Hills, Balochistan, Pakistan. Holotype (ISEM DBJ2−A1). A. Stereophoto of occlusal view. B. Stereophoto of labial view. Scale bars 1 cm.

écaille Pakistan Chattien Oligocène +1
Holotype skull of the centrosaurine ceratopsian dinosaur Diabloceratops eatoni from the Late Cretaceous (Campanian) of the Wahweap Formation, Grand Staircase-Escalante National Monument, Utah. Photograph by Nick Longrich, 2023.

Holotype skull of the centrosaurine ceratopsian dinosaur Diabloceratops eatoni from the Late Cretaceous (Campanian) of the Wahweap Formation, Grand Staircase-Escalante National Monument, Utah. Photograph by Nick Longrich, 2023.

Campanien Crétacé Crétacé supérieur holotype +5
Arisierpeton simplex, holotype specimen premaxilla

Arisierpeton simplex, holotype specimen premaxilla

holotype spécimen
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