holotype

Specimen nature

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Positive digital cast of the holotype of the pterosaur Herbstosaurus pigmaeus (CTES-PZ 1711). Abbreviations: gtr, greater trochanter; isp, ischial peduncle; l.fe, left femur; ppush, prepubic shaft; prap, preacetabular process; r.fe, right femur; r.il, right ilium; r.ppu, right prepubis; sr5, sacral rib 5; sv1, sacral vertebra 1. Scale bar equals 2 cm.
Taxa Herbstosaurus

Positive digital cast of the holotype of the pterosaur Herbstosaurus pigmaeus (CTES-PZ 1711). Abbreviations: gtr, greater trochanter; isp, ischial peduncle; l.fe, left femur; ppush, prepubic shaft; prap, preacetabular process; r.fe, right femur; r.il, right ilium; r.ppu, right prepubis; sr5, sacral rib 5; sv1, sacral vertebra 1. Scale bar equals 2 cm.

scale vertebra cast holotype +2
Figure 2.
Holotype of Liaodactylus primus gen. et sp. nov. (PMOL-AP00031): photographs and line drawings of the nearly complete skull with mandibles in left lateral (a,b), and palatal (c,d) views. aa, atlas-axis complex; an, angular; arf, articular facet; bo, basioccipital; bs, basisphenoid; co, coronoid; d, dentary; ec, ectopterygoid; f, frontal; j, jugal; m, maxilla; naof, nasoantorbital fenestra; p, parietal; pal, palatine; pat, proatlas; pecf, pterygo-ectopterygoid fenestra; pm, premaxilla; pof, postfrontal; pra, prearticular; pt, pterygoid; q, quadrate; qj, quadratojugal; sa, surangular; sof, suborbital fenestra; spl, splenial; sq, squamosal; stf, subtemporal fenestra.
Taxa Liaodactylus

Figure 2. Holotype of Liaodactylus primus gen. et sp. nov. (PMOL-AP00031): photographs and line drawings of the nearly complete skull with mandibles in left lateral (a,b), and palatal (c,d) views. aa, atlas-axis complex; an, angular; arf, articular facet; bo, basioccipital; bs, basisphenoid; co, coronoid; d, dentary; ec, ectopterygoid; f, frontal; j, jugal; m, maxilla; naof, nasoantorbital fenestra; p, parietal; pal, palatine; pat, proatlas; pecf, pterygo-ectopterygoid fenestra; pm, premaxilla; pof, postfrontal; pra, prearticular; pt, pterygoid; q, quadrate; qj, quadratojugal; sa, surangular; sof, suborbital fenestra; spl, splenial; sq, squamosal; stf, subtemporal fenestra.

drawing holotype Liaodactylus skull
Pterodactylus sagittirostris, holotype NHMUK PV R 1823 (upper Berriasian / Valangianian, Hastings Group), part of the mandibular rami. A right lateral view B respective line drawing C left lateral view D respective line drawing E ventral view. Abbreviation: d – dentary. Arrows indicate alveoli or teeth. Scale bar = 10 mm. Photos courtesy of The Natural History Museum.
Taxa Serradraco

Pterodactylus sagittirostris, holotype NHMUK PV R 1823 (upper Berriasian / Valangianian, Hastings Group), part of the mandibular rami. A right lateral view B respective line drawing C left lateral view D respective line drawing E ventral view. Abbreviation: d – dentary. Arrows indicate alveoli or teeth. Scale bar = 10 mm. Photos courtesy of The Natural History Museum.

scale drawing museum Berriasian +3
Holotype of Eosipterus yangi GMV 2117, displaying soft tissue preservation around the hind limbs and/or ankles.
Taxa Eosipterus

Holotype of Eosipterus yangi GMV 2117, displaying soft tissue preservation around the hind limbs and/or ankles.

limb tissue holotype Eosipterus
FSAC-OB 7, holotype right humerus, Simurghia robustus.
In (A), dorsal view, (B) ventral view, and (C) posterior view. Abbreviations: dpc, deltopectoral crest; msc, muscle scar; scpr, supracondylar process of the ectepicondyle; vp, ventral pillar; vt, ventral tubercle of the deltopectoral crest.
Taxa Simurghia

FSAC-OB 7, holotype right humerus, Simurghia robustus. In (A), dorsal view, (B) ventral view, and (C) posterior view. Abbreviations: dpc, deltopectoral crest; msc, muscle scar; scpr, supracondylar process of the ectepicondyle; vp, ventral pillar; vt, ventral tubercle of the deltopectoral crest.

crest humerus holotype Simurghia
FSAC-OB 232, holotype skeleton of Barbaridactylus grandis.
(A) Left humerus in anterior view, (B) right femur in anterior view, (C) right radius and ulna in posterior view, (D) cervical vertebra in ventral view, (E) left scapulocoracoid in medial view, and (F) posterior ramus of the right mandible in medial view.

Abbreviations: co, coracoid; cot, cotyle of mandible; cvr, cervical ribs; dm, dorsal margin of mandible; dpc, deltopectoral crest; gl, glenoid; fh, femoral head; gt, greater trochanter; hh, humeral head; hyp, hypapophysis; olp, olecranon process; pex, postexapophysis; pn, pneumatopore; rad, radius; sca, scapula; scp, supracondylar process of the ectepicondyle; uc, ulnar crest; uln, ulna; vm, ventral margin of mandible.
Taxa Barbaridactylus

FSAC-OB 232, holotype skeleton of Barbaridactylus grandis. (A) Left humerus in anterior view, (B) right femur in anterior view, (C) right radius and ulna in posterior view, (D) cervical vertebra in ventral view, (E) left scapulocoracoid in medial view, and (F) posterior ramus of the right mandible in medial view. Abbreviations: co, coracoid; cot, cotyle of mandible; cvr, cervical ribs; dm, dorsal margin of mandible; dpc, deltopectoral crest; gl, glenoid; fh, femoral head; gt, greater trochanter; hh, humeral head; hyp, hypapophysis; olp, olecranon process; pex, postexapophysis; pn, pneumatopore; rad, radius; sca, scapula; scp, supracondylar process of the ectepicondyle; uc, ulnar crest; uln, ulna; vm, ventral margin of mandible.

crest humerus vertebra holotype +2
Epapatelo otyikokolo holotype (MGUAN-PA650), left ulna. A) proximal,
(B) posterior, (C) distal, (D) dorsal, (E) anterior, and (F) ventral views. Abbreviations: AC, anterior cotyle; B, bump; BT, biceps tubercle; F, fovea; FTG, flexor tendon groove; MS, muscle scar; OP, olecranon process; PC, posterior cotyle; PF, pneumatic foramen; and UT, ulnar tubercle; IMS, interosseous membrane scar; VP, ventral process.
Taxa Epapatelo

Epapatelo otyikokolo holotype (MGUAN-PA650), left ulna. A) proximal, (B) posterior, (C) distal, (D) dorsal, (E) anterior, and (F) ventral views. Abbreviations: AC, anterior cotyle; B, bump; BT, biceps tubercle; F, fovea; FTG, flexor tendon groove; MS, muscle scar; OP, olecranon process; PC, posterior cotyle; PF, pneumatic foramen; and UT, ulnar tubercle; IMS, interosseous membrane scar; VP, ventral process.

holotype Epapatelo
Holotype of Haopterus gracilis on display at the Paleozoological Museum of China.
Taxa Haopterus

Holotype of Haopterus gracilis on display at the Paleozoological Museum of China.

museum China holotype Haopterus +1
Holotype skull (IVPP V17083) of Guidraco venator on display at the Paleozoological Museum of China.
Taxa Guidraco

Holotype skull (IVPP V17083) of Guidraco venator on display at the Paleozoological Museum of China.

museum China holotype Guidraco +1
Holotype skull and mandible of the anhanguerid Ferrodraco (A), holotype skull and mandible of the anhanguerid Mythunga (B), and holotype mandible of the targaryendraconian Aussiedraco (C).
Taxa Mythunga

Holotype skull and mandible of the anhanguerid Ferrodraco (A), holotype skull and mandible of the anhanguerid Mythunga (B), and holotype mandible of the targaryendraconian Aussiedraco (C).

holotype Anhangueridae Aussiedraco Ferrodraco +3
Ferrodraco lentoni gen. et sp. nov. holotype specimen AODF 876. All preserved elements were photographed and scaled to the same size, then articulated where possible. These were then used as the basis for the scaling of the skeletal reconstruction, the missing parts of which were based on the skeletal reconstruction of Tropeognathus mesembrinus by Mark Witton. Scale bar = 50 mm.
Taxa Ferrodraco

Ferrodraco lentoni gen. et sp. nov. holotype specimen AODF 876. All preserved elements were photographed and scaled to the same size, then articulated where possible. These were then used as the basis for the scaling of the skeletal reconstruction, the missing parts of which were based on the skeletal reconstruction of Tropeognathus mesembrinus by Mark Witton. Scale bar = 50 mm.

scale holotype specimen Ferrodraco +1
Haliskia peterseni gen. et sp. nov., holotype specimen KK F1426. (A) dorsal vertebra, ribs, gastralium, left scapulocoracoid, left and right manual phalanx IV-2, right manual phalanx IV-3, left femur, left tibia, metatarsals and pedal phalanges, (B) mandible, teeth, ceratobranchials, cervical vertebra, rib, gastralium, right syncarpus, right lateral carpal, right pteroid, metacarpals, manual digits, right manual phalanx IV-1, left manual phalanx IV-4, (C) premaxilla with teeth, (D) isolated tooth, (E) isolated tooth, (F) left lateral carpal, (G) left metacarpal IV. Scale bar = 50 mm.
Taxa Haliskia

Haliskia peterseni gen. et sp. nov., holotype specimen KK F1426. (A) dorsal vertebra, ribs, gastralium, left scapulocoracoid, left and right manual phalanx IV-2, right manual phalanx IV-3, left femur, left tibia, metatarsals and pedal phalanges, (B) mandible, teeth, ceratobranchials, cervical vertebra, rib, gastralium, right syncarpus, right lateral carpal, right pteroid, metacarpals, manual digits, right manual phalanx IV-1, left manual phalanx IV-4, (C) premaxilla with teeth, (D) isolated tooth, (E) isolated tooth, (F) left lateral carpal, (G) left metacarpal IV. Scale bar = 50 mm.

scale tooth vertebra holotype +2
Skull of Caiuajara dobruskii gen. et sp. nov. (holotype, CP.V 1449) with the shape of an adult individual.

Scale bar equals 50: d, dentary; dcr, dentary crest; dep, depression; exp, ventral expansion of the premaxilla; f, frontal; fcr, frontal crest; fo, foraminae; m, maxilla; oc, occipital condyle; op, opisthotic; p, parietal; pm, premaxilla; pmcr, premaxillary crest; q, quadrate; soc, supraoccipital. The quadrate is inverted.
Taxa Caiuajara

Skull of Caiuajara dobruskii gen. et sp. nov. (holotype, CP.V 1449) with the shape of an adult individual. Scale bar equals 50: d, dentary; dcr, dentary crest; dep, depression; exp, ventral expansion of the premaxilla; f, frontal; fcr, frontal crest; fo, foraminae; m, maxilla; oc, occipital condyle; op, opisthotic; p, parietal; pm, premaxilla; pmcr, premaxillary crest; q, quadrate; soc, supraoccipital. The quadrate is inverted.

crest scale holotype Caiuajara +1
Holotype of Aymberedactylus cearensis gen. et sp. nov. (MN 7596-V).

(A) Dorsal view. (B) Ventral view. (C) Right lateral view. Abbreviations: ang = angular, d = dentary, dfo = dentary fossa, glfo = glenoid fossa, rapr = retroarticular process, san = surangular, sys = symphyseal shelf.
Taxa Aymberedactylus

Holotype of Aymberedactylus cearensis gen. et sp. nov. (MN 7596-V). (A) Dorsal view. (B) Ventral view. (C) Right lateral view. Abbreviations: ang = angular, d = dentary, dfo = dentary fossa, glfo = glenoid fossa, rapr = retroarticular process, san = surangular, sys = symphyseal shelf.

holotype Aymberedactylus
Azhdarchoid pterosaur Kariridraco dianae gen. et sp. nov. holotype (MPSC R 1056) from Lower Cretaceous of Northeastern Brazil. Skull in right lateral view. Photograph (A1), interpretative drawing (A2). Dark grey areas indicate remains of the carbonate matrix still attachaed to the skull.
Taxa Kariridraco

Azhdarchoid pterosaur Kariridraco dianae gen. et sp. nov. holotype (MPSC R 1056) from Lower Cretaceous of Northeastern Brazil. Skull in right lateral view. Photograph (A1), interpretative drawing (A2). Dark grey areas indicate remains of the carbonate matrix still attachaed to the skull.

drawing Brazil Cretaceous holotype +3
The holotype of the pterosaur Lacusovagus
Taxa Lacusovagus

The holotype of the pterosaur Lacusovagus

holotype Lacusovagus Pterosauria
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