holotype

Dinosaur National Monument is a United States National Monument located on the southeast flank of the Uinta Mountains on the border between Colorado and Utah at the confluence of the Green and Yampa Rivers. Although most of the monument area is in Moffat County, Colorado, the Dinosaur Quarry is located in Utah just to the north of the town of Jensen, Utah. The nearest communities are Jensen, Utah, and Dinosaur, Colorado. The park contains over 800 paleontological sites and has fossils of dinosaurs including Allosaurus, Deinonychus, Abydosaurus (a nearly complete skull, lower jaws and first four neck vertebrae of the specimen DINO 16488 found here at the base of the Mussentuchit Member of the Cedar Mountain Formation is the holotype for the description) and various long-neck, long-tail sauropods. It was declared a National Monument on October 4, 1915. The rock layer enclosing the fossils is a sandstone and conglomerate bed of alluvial or river bed origin known as the Morrison Formation from the Jurassic Period some 150 million years old. The dinosaurs and other ancient animals were carried by the river system which eventually entombed their remains in Utah. The pile of sediments were later buried and lithified into solid rock. The layers of rock were later uplifted and tilted to their present angle by the mountain building forces that formed the Uintas during the Laramide orogeny. The relentless forces of erosion exposed the layers at the surface to be found by paleontologists. The dinosaur fossil beds (bone beds) were discovered in 1909 by Earl Douglass, a paleontologist working and collecting for the Carnegie Museum of Natural History. He and his crews excavated thousands of fossils and shipped them back to the museum in Pittsburgh, Pennsylvania for study and display. President Woodrow Wilson proclaimed the dinosaur beds as Dinosaur National Monument in 1915. The monument boundaries were expanded in 1938 from the original 80-acre (320,000 m2) tract surrounding the dinosaur quarry in Utah, to its present extent of over 200,000 acres (800 km²) in Utah and Colorado, encompassing the spectacular river canyons of the Green and Yampa. Though lesser-known than the fossil beds, the petroglyphs in Dinosaur National Monument are another treasure the monument holds. Due to problems with vandals, many of the sites are not listed on area maps. The "Wall of Bones" located within the Dinosaur Quarry building in the park consists of a steeply tilted (67° from horizontal) rock layer which contains hundreds of dinosaur fossils. The enclosing rock has been chipped away to reveal the fossil bones intact for public viewing. In July 2006, the Quarry Visitor Center was closed due to structural problems that since 1957 had plagued the building because it was built on unstable clay. The decision was made to build a new facility elsewhere in the monument to house the visitor center and administrative functions, making it easier to resolve the structural problems of the quarry building while still retaining a portion of the historic Mission 66 era exhibit hall. It was announced in April 2009 that Dinosaur National Monument would receive $13.1 million to refurbish and reopen the gallery as part of the Obama administration's $750 billion stimulus plan. The Park Service successfully rebuilt the Quarry Exhibit Hall, supporting its weight on 70-foot steel micropile columns that extend to the bedrock below the unstable clay. The Dinosaur Quarry was reopened in Fall 2011. en.wikipedia.org/wiki/Dinosaur_National_Monument en.wikipedia.org/wiki/Wikipedia:Text_of_Creative_Commons_...

Life restoration of the Triassic ichthyosaur Callawayia neoscapularis. Three specimens of this ichthyosaur are known, the holotype, ROM 41993, and two referred specimens, TMP 94.380.11 and 94.382.2. The skull is primarily based on ROM 41993, cross-checked against TMP 94.380.11 and TMP 94.382.2. The vertebral column is based primarily on TMP 94.382.2 as it is the most complete of these specimens, while the ribs were based on ROM 41993. The forelimbs were mainly based on those of ROM 41993, with TMP 94.380.11 used to determine their breadth. The hindlimbs were based on TMP 94.380.11, especially the more complete right hindlimb. ROM 41993 was cross-scaled with TMP 94.380.11 by the dimensions of the forelimb epipodials, which produced similar vertebral dimensions. The two TMP specimens were cross-scaled based on femoral length, also producing similar vertebral dimensions. Nicholls & Manabe (2001) stated that no wedge-shaped caudal centra supporting a tailbend were found and that there was no evidence of a bend being present, though considered that they might have existed in the gap in the preserved caudals. Since various other Triassic ichthyosaurs have since been found to have tail bends, one was illustrated here. A modest downturn of roughly 15° was illustrated, comparable to that in Guanlingsaurus, and the location of the bend within the gap in the preserved vertebrae matches well with the location of the bend in Guizhouichthyosaurus. References McGowan, C. (1994). "A new species of Shastasaurus (Reptilia: Ichthyosauria) from the Triassic of British Columbia: The most complete exemplar of the genus". Journal of Vertebrate Paleontology 14 (2): 168–179. DOI:10.1080/02724634.1994.10011550. Nicholls, E. L.; Manabe, M. (2001). "A new genus of ichthyosaur from the Late Triassic Pardonet Formation of British Columbia: Bridging the Triassic-Jurassic gap". Canadian Journal of Earth Sciences 38 (6): 983–1002. Ji, C.; Jiang, D.Y.; Hao, W.; Sun, Y. (2011). "True tailbend occurred in the Late Triassic: Evidence from ichthyosaur skeletons of South China". Acta Scientiarum Naturalium Universitatis Pekinensis 47 (2): 309–314. Shang, Q. H.; Li, C. (2009). "On the occurrence of the ichthyosaur Shastasaurus in the Guanling biota (Late Triassic), Guizhou, China". Vertebrata PalAsiatica 47 (3): 178–193.

Fostoria dhimbangunmal (3D digital rendering of LRF 3050; holotype) braincase in I, dorsal and J, ventral views. Scale bar = 10 cm.

Shetwemys, Shell remains of the podocnemidid turtle S. fajumensis (Erymnochelyini). (a–c) SMNS 11233/2, partial carapace, in dorsal (a), ventral (b), and left lateral (c) views. (d) Ventral view of the anterior lobe the holotype of the species, currently lost, based on the fig. 2C in plate 8 of Andrews (1903). (e–g) SMNS 12647, plastron, in ventral (e), dorsal (f), and left lateral (g) views. (g’) corresponds to an enlarged photograph of the posterior plastral lobe, in left lateral view, in which the thickness in the regions close to the hypo-xiphiplastral suture (in blue), between the pelvic scars (in green), and at the level of the anal notch (in red), have been represented by arrows (h–i), SMNS 12646, plastron, in ventral (h) and dorsal (i) views. Gebel Quatrani Formation, Fayum depression, Egypt, Lower Oligocene (Rupelian)

Close up of the Eulithomyrmex rugosus holotype head. Museum of Comparative Zoology specimen UCM17019. Priabonian; Florissant Formation, Colorado, USA

The Holotype batomorph tooth of Antiquaobatis grimmenensis from the Late Pliensbachian (spinatum) of Grimmen (Mecklenburg-Western Pomerania, Germany).

New species of Mesozoic benthic foraminifera from the former British Petroleum micropalaeontology collection, Ophthalmidium dracomaris n. sp. (1) holotype (NHMUK PM PF 74496); (2–5) paratypes (NHMUK PM PF 74497–74500). (8–9) Eobigenerina calloviensis n. sp.: (8) holotype (NHMUK PM PF 74504); (9) paratype (NHMUK PM PF 74505). (10–12) Trochammina fordonensis n. sp.: (10) holotype (NHMUK PM PF 74501); (11–12) paratype (NHMUK PM PF 74502–74503). (13–15) Arenoturrispirillina swiecickii n. sp.: (13) holotype (NHMUK PM PF 74506); (14–15) paratype (NHMUK PM PF 74507 + NHMUK PM PF 74552). (16–17) Ataxophragmium mariae n. sp.: (16) holotype (NHMUK PM PF 74553); (17) paratype (NHMUK PM PF 74554).

Eekaulostomus cuevasae gen. and sp. nov. from the Danian marine deposits of the Belisario Domínguez quarry, near Palenque, Chiapas, southeastern Mexico. 1, General view of IGM 4716, holotype and single specimen known. 2, Idealized line drawing of the same specimen.

Palaeohypsodontus zinensis sp. nov., late Oligocene (Chattian), Lundo J2, Bugti Hills, Balochistan, Pakistan. Holotype (ISEM DBJ2−A1). A. Stereophoto of occlusal view. B. Stereophoto of labial view. Scale bars 1 cm.

Holotype skull of the centrosaurine ceratopsian dinosaur Diabloceratops eatoni from the Late Cretaceous (Campanian) of the Wahweap Formation, Grand Staircase-Escalante National Monument, Utah. Photograph by Nick Longrich, 2023.

Arisierpeton simplex, holotype specimen premaxilla

Fossil of Mei long (holotype), on temporary display at the Shanghai Natural History Museum as part of the exhibition "China's Dinosaur World". Shot on July 22, 2025.

Holotype block of Oksoko avarsan MPC-D 102/110. (a,b) Holotype block with skeletons in ventral view. Colours distinguish different individuals; the holotype individual is in blue.

Fossil samples – e.g. ‘primitive’ bony fish (1, 2), a skull of a temnospondyl ‘amphibian’ (probably a metoposauroid) in dorsal view (3), a skull of an archosaur of the crocodile lineage (probably a phytosaur) in palatal view (4), holotype of the “gliding reptile” Icarosaurus siefkeri [1] (5) and Atreipus-Grallator-type dinosaur tracks (bottom right) – from the Newark Supergroup, i.e. a series of mainly Late Triassic to Early Jurassic sedimentary rocks of eastern North America ↑ Edwin H. Colbert: A gliding reptile from the Triassic of New Jersey. American Museum Novitates, 2230. American Museum of Natural History, New York 1966, digitallibrary.amnh.org, cf. fig. 3 therein.

Carinodens acrodon holotype left maxilla (MHNM.KHG.1510) (A) compared to Xenodens calminechari holotype left maxilla (MHNM.KHG.331) (B), in lateral view; both from Sidi Chennane, Oulad Abdoun Basin, Morocco; Phosphates, Upper Couche III, uppermost Maastrichtian [9,24]. Scales = 50 mm.

Holotype of Jian changmaensis (GSGM-D050), an articulated partial left pectoral girdle (scapulocoracoid) and forelimb (humerus, radius, and ulna). A, silhouette of generalized microraptorine dromaeosaurid theropod (courtesy Scott Hartman) showing skeletal elements preserved; B, photograph of specimen as preserved, exposed primarily in dorsomedial (scapulocoracoid), caudodorsal (humerus), and dorsal (radius and ulna) views; C, interpretive line drawing of B; D, detail photograph of scapulocoracoid and proximal end of humerus in caudodorsal view, showing supracoracoid fenestra and other structures; E, interpretive line drawing of D. Abbreviations: ac, acromion; bc, bicipital crest; C, coracoid; cr, caudal ridge; dep, dorsal epicondyle; dpc, deltopectoral crest; dr, dorsal ridge; ed, epicondylar depression; fs?, fossa for M. supinator?; H, humerus; hh, humeral head; lp, lateral process; ‘mb’, ‘medial bar’; op, olecranon process; R, radius; S, scapula; scb, scapular blade; scf, supracoracoid fenestra; sta, sternal articulation; U, ulna.