holotype

Nature du spécimen

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Fossil of Mei long (holotype), on temporary display at the Shanghai Natural History Museum as part of the exhibition "China's Dinosaur World". Shot on July 22, 2025.

Fossil of Mei long (holotype), on temporary display at the Shanghai Natural History Museum as part of the exhibition "China's Dinosaur World". Shot on July 22, 2025.

musée Chine fossile holotype +1
Holotype block of Oksoko avarsan MPC-D 102/110. (a,b) Holotype block with skeletons in ventral view. Colours distinguish different individuals; the holotype individual is in blue.

Holotype block of Oksoko avarsan MPC-D 102/110. (a,b) Holotype block with skeletons in ventral view. Colours distinguish different individuals; the holotype individual is in blue.

holotype Oksoko squelette
Fossil samples – e.g. ‘primitive’ bony fish (1, 2), a skull of a temnospondyl ‘amphibian’ (probably a metoposauroid) in dorsal view (3), a skull of an archosaur of the crocodile lineage (probably a phytosaur) in palatal view (4), holotype of the “gliding reptile” Icarosaurus siefkeri [1] (5) and Atreipus-Grallator-type dinosaur tracks (bottom right) – from the Newark Supergroup, i.e. a series of mainly Late Triassic to Early Jurassic sedimentary rocks of eastern North America


↑ Edwin H. Colbert: A gliding reptile from the Triassic of New Jersey. American Museum Novitates, 2230. American Museum of Natural History, New York 1966, digitallibrary.amnh.org, cf. fig. 3 therein.

Fossil samples – e.g. ‘primitive’ bony fish (1, 2), a skull of a temnospondyl ‘amphibian’ (probably a metoposauroid) in dorsal view (3), a skull of an archosaur of the crocodile lineage (probably a phytosaur) in palatal view (4), holotype of the “gliding reptile” Icarosaurus siefkeri [1] (5) and Atreipus-Grallator-type dinosaur tracks (bottom right) – from the Newark Supergroup, i.e. a series of mainly Late Triassic to Early Jurassic sedimentary rocks of eastern North America ↑ Edwin H. Colbert: A gliding reptile from the Triassic of New Jersey. American Museum Novitates, 2230. American Museum of Natural History, New York 1966, digitallibrary.amnh.org, cf. fig. 3 therein.

musée Jurassique inférieur Jurassique Trias supérieur +8
Carinodens acrodon holotype left maxilla (MHNM.KHG.1510) (A) compared to Xenodens calminechari holotype left maxilla (MHNM.KHG.331) (B), in lateral view; both from Sidi Chennane, Oulad Abdoun Basin, Morocco; Phosphates, Upper Couche III, uppermost Maastrichtian [9,24]. Scales = 50 mm.

Carinodens acrodon holotype left maxilla (MHNM.KHG.1510) (A) compared to Xenodens calminechari holotype left maxilla (MHNM.KHG.331) (B), in lateral view; both from Sidi Chennane, Oulad Abdoun Basin, Morocco; Phosphates, Upper Couche III, uppermost Maastrichtian [9,24]. Scales = 50 mm.

écaille Maroc Maastrichtien holotype +2
Holotype of Jian changmaensis (GSGM-D050), an articulated partial left pectoral girdle (scapulocoracoid) and forelimb (humerus, radius, and ulna). A, silhouette of generalized microraptorine dromaeosaurid theropod (courtesy Scott Hartman) showing skeletal elements preserved; B, photograph of specimen as preserved, exposed primarily in dorsomedial (scapulocoracoid), caudodorsal (humerus), and dorsal (radius and ulna) views; C, interpretive line drawing of B; D, detail photograph of scapulocoracoid and proximal end of humerus in caudodorsal view, showing supracoracoid fenestra and other structures; E, interpretive line drawing of D. Abbreviations: ac, acromion; bc, bicipital crest; C, coracoid; cr, caudal ridge; dep, dorsal epicondyle; dpc, deltopectoral crest; dr, dorsal ridge; ed, epicondylar depression; fs?, fossa for M. supinator?; H, humerus; hh, humeral head; lp, lateral process; ‘mb’, ‘medial bar’; op, olecranon process; R, radius; S, scapula; scb, scapular blade; scf, supracoracoid fenestra; sta, sternal articulation; U, ulna.

Holotype of Jian changmaensis (GSGM-D050), an articulated partial left pectoral girdle (scapulocoracoid) and forelimb (humerus, radius, and ulna). A, silhouette of generalized microraptorine dromaeosaurid theropod (courtesy Scott Hartman) showing skeletal elements preserved; B, photograph of specimen as preserved, exposed primarily in dorsomedial (scapulocoracoid), caudodorsal (humerus), and dorsal (radius and ulna) views; C, interpretive line drawing of B; D, detail photograph of scapulocoracoid and proximal end of humerus in caudodorsal view, showing supracoracoid fenestra and other structures; E, interpretive line drawing of D. Abbreviations: ac, acromion; bc, bicipital crest; C, coracoid; cr, caudal ridge; dep, dorsal epicondyle; dpc, deltopectoral crest; dr, dorsal ridge; ed, epicondylar depression; fs?, fossa for M. supinator?; H, humerus; hh, humeral head; lp, lateral process; ‘mb’, ‘medial bar’; op, olecranon process; R, radius; S, scapula; scb, scapular blade; scf, supracoracoid fenestra; sta, sternal articulation; U, ulna.

crête humérus dessin holotype +3
Holotype of Rexarthuria or "Palaeornis" clifti, PV OR 2353. Proximal extremity of left humerus.
Taxons Rexarthuria

Holotype of Rexarthuria or "Palaeornis" clifti, PV OR 2353. Proximal extremity of left humerus.

humérus musée holotype Palaeornis +1
Representative taxa from the Santonian Iharkút fauna from the Csehbánya Formation, Bakony Mountains, western Hungary. A Pannoniasaurus inexpectatus (Squamata, Mosasauroidea), dorsal vertebra (MTM uncatalogued) in dorsal view (photo by Réka Kalmár) B Foxemys trabanti (Pleurodira, Bothremydidae), skull (MTM V 2010.215.1.) in dorsal view (photo by Márton Rabi). C Bicuspidon aff. hatzegiensis (Squamata, Borioteiioidea), left dentary (MTM 2006.112.1.) in medial view (photo by László Makádi) D Basal tetanuran (Theropoda, Tetanurae), tooth (MTM V.01.54) in ?lingual view E Indeterminate abelisaurid (Theropoda, Abelisauridae), pedal ungual phalanx (MTM V 2008.43.1.) in lateral view F Pneumatoraptor fodori (Theropoda, Paraves), left scapulocoracoid (holotype, MTM V 2008.38.1.) in lateral view G Mochlodon vorosi (Ornithopoda, Rhabdodontidae), left dentary (holotype, MTM V 2010.105.1) in lateral view H Bakonydraco galaczi (Pterosauria, Azhdarchidae), mandible (holotype, MTM 2007.110.1) in dorsal view I Iharkutosuchus makadii (Eusuchia, Hylaeochampsidae), skull (holotype, MTM 2006.52.1) in dorsal view J Hungarosaurus tormai (Ankylosauria, Nodosauridae), right dentary (MTM 2007.25.2) in lateral view K Bauxitornis mindszentyae (Aves, Enantiornithes), left tarsometatarsus (holotype, MTM V 2009.38.1) in anterior view L Ajkaceratops kozmai (Ceratopsia), fused rostral and premaxillae (holotype, MTM V 2009.192.1) in lateral view. Scale bars: 2 cm in A, V, G, H, I, J; 1 cm in D, E, F, K, L; 1 mm in C.

Representative taxa from the Santonian Iharkút fauna from the Csehbánya Formation, Bakony Mountains, western Hungary. A Pannoniasaurus inexpectatus (Squamata, Mosasauroidea), dorsal vertebra (MTM uncatalogued) in dorsal view (photo by Réka Kalmár) B Foxemys trabanti (Pleurodira, Bothremydidae), skull (MTM V 2010.215.1.) in dorsal view (photo by Márton Rabi). C Bicuspidon aff. hatzegiensis (Squamata, Borioteiioidea), left dentary (MTM 2006.112.1.) in medial view (photo by László Makádi) D Basal tetanuran (Theropoda, Tetanurae), tooth (MTM V.01.54) in ?lingual view E Indeterminate abelisaurid (Theropoda, Abelisauridae), pedal ungual phalanx (MTM V 2008.43.1.) in lateral view F Pneumatoraptor fodori (Theropoda, Paraves), left scapulocoracoid (holotype, MTM V 2008.38.1.) in lateral view G Mochlodon vorosi (Ornithopoda, Rhabdodontidae), left dentary (holotype, MTM V 2010.105.1) in lateral view H Bakonydraco galaczi (Pterosauria, Azhdarchidae), mandible (holotype, MTM 2007.110.1) in dorsal view I Iharkutosuchus makadii (Eusuchia, Hylaeochampsidae), skull (holotype, MTM 2006.52.1) in dorsal view J Hungarosaurus tormai (Ankylosauria, Nodosauridae), right dentary (MTM 2007.25.2) in lateral view K Bauxitornis mindszentyae (Aves, Enantiornithes), left tarsometatarsus (holotype, MTM V 2009.38.1) in anterior view L Ajkaceratops kozmai (Ceratopsia), fused rostral and premaxillae (holotype, MTM V 2009.192.1) in lateral view. Scale bars: 2 cm in A, V, G, H, I, J; 1 cm in D, E, F, K, L; 1 mm in C.

dent vertèbre Hongrie Santonien +15
Anhanguera blittersdorffi holotype (MN 4805-V) in lateral view.

Anhanguera blittersdorffi holotype (MN 4805-V) in lateral view.

holotype Anhanguera
Complete specimen of the Laueropterus vitriolus holotype (LF 6268). The holotype specimen seen under natural light. Major elements are labeled. Abbreviations as follows here and in subsequent figures: ca, caudal vertebra; cp, carpal; cv, cervical vertebrae; dv, dorsal vertebra; fe, femur; hu, humerus; mn, mandible; mt, metatarsal; ph, phalanges; pmm, premaxilla-maxilla; pp, prepubis; pt, pteroid; pv, pelvic plate; r, rib; sa, sacrum; sc, scapulocoracoid; sk, skull part; st, sternum; ta, tarsals; ti, tibia; ul, ulna; un, ungual; wpx, wing phalanges; wmc, wing metacarpal. Scale bar = 100 mm.
Taxons Laueropterus

Complete specimen of the Laueropterus vitriolus holotype (LF 6268). The holotype specimen seen under natural light. Major elements are labeled. Abbreviations as follows here and in subsequent figures: ca, caudal vertebra; cp, carpal; cv, cervical vertebrae; dv, dorsal vertebra; fe, femur; hu, humerus; mn, mandible; mt, metatarsal; ph, phalanges; pmm, premaxilla-maxilla; pp, prepubis; pt, pteroid; pv, pelvic plate; r, rib; sa, sacrum; sc, scapulocoracoid; sk, skull part; st, sternum; ta, tarsals; ti, tibia; ul, ulna; un, ungual; wpx, wing phalanges; wmc, wing metacarpal. Scale bar = 100 mm.

humérus écaille vertèbre holotype +3
Cryptarcus ("Chasmosaurus") russelli holotype cranium (CMNFV 8800). (A) Skull in left lateral view (with restored jugal and lower jaw); (B) detail of snout in left lateral view; (C) detail of parietosquamosal frill in dorsal view; (D) detail of supraorbital review in dorsal view; (E) detail of es6 episquamosal; (F) detail of es7 episquamosal. Arrowheads in E and F point to margins of episquamosals.
Taxons Cryptarcus

Cryptarcus ("Chasmosaurus") russelli holotype cranium (CMNFV 8800). (A) Skull in left lateral view (with restored jugal and lower jaw); (B) detail of snout in left lateral view; (C) detail of parietosquamosal frill in dorsal view; (D) detail of supraorbital review in dorsal view; (E) detail of es6 episquamosal; (F) detail of es7 episquamosal. Arrowheads in E and F point to margins of episquamosals.

holotype Chasmosaurus Cryptarcus crâne
The holotype and only known specimen of the hauffiopterygian leptonectid, Xiphodracon goldencapensis (ROM VP52596) from Golden Cap, between Charmouth and Seatown, Dorset, UK. The skeleton is exposed in ventrolateral view. The skull has been fully prepared free of matrix whereas most of the skeleton is still in matrix. The left (upper) forefin has been prepared so that it is three-dimensionally preserved and projects upwards. Scale bar represents 20 cm.

The holotype and only known specimen of the hauffiopterygian leptonectid, Xiphodracon goldencapensis (ROM VP52596) from Golden Cap, between Charmouth and Seatown, Dorset, UK. The skeleton is exposed in ventrolateral view. The skull has been fully prepared free of matrix whereas most of the skeleton is still in matrix. The left (upper) forefin has been prepared so that it is three-dimensionally preserved and projects upwards. Scale bar represents 20 cm.

écaille Royaume-Uni holotype spécimen +5
Oneirosaurus caballeroi holotype IGMp879524. A, photograph, drawing and interpretative scheme of the specimen without left mandibular ramus, in right lateral view. B, photograph, drawing and interpretative scheme of the specimen without left mandibular
ramus, in left lateral view. Gray fill, sediment; pattern fill, broken bone surface; black, empty space. Scale bars: A = 50 mm; C-D = 30 mm. Abbreviations: an, angular; avc, anterior aperture of the vidian canal; bo, basioccipital; bptm, basipterygoid meniscus; bs, basisphenoid; c, coronoid; d, dentary; ec, ectopterygoid; ep, epipterygoid; ex, exoccipital; f, frontal; im, internal auditory meatus; is, interorbital septum; j, jugal; la, lacrimal; m, mandible; mx, maxilla; na, nasal; op, opisthotic-exoccipital; p, parietal; par, prearticular; pl, palatine; pmx, premaxilla; pof, postorbitofrontal; pr, prootic; prf, prefrontal; ps, parasphenoid; pt, pterygoid; pvc, posterior aperture of the vidian canal q; quadrate; rv, right vomer; sa, surangular; scp, sclerotic plate; smx, septomaxilla; so, supraoccipital; sp, splenial; stp, stapes; VII, IX-XII, exit of cranial nerves.
Taxons Oneirosaurus

Oneirosaurus caballeroi holotype IGMp879524. A, photograph, drawing and interpretative scheme of the specimen without left mandibular ramus, in right lateral view. B, photograph, drawing and interpretative scheme of the specimen without left mandibular ramus, in left lateral view. Gray fill, sediment; pattern fill, broken bone surface; black, empty space. Scale bars: A = 50 mm; C-D = 30 mm. Abbreviations: an, angular; avc, anterior aperture of the vidian canal; bo, basioccipital; bptm, basipterygoid meniscus; bs, basisphenoid; c, coronoid; d, dentary; ec, ectopterygoid; ep, epipterygoid; ex, exoccipital; f, frontal; im, internal auditory meatus; is, interorbital septum; j, jugal; la, lacrimal; m, mandible; mx, maxilla; na, nasal; op, opisthotic-exoccipital; p, parietal; par, prearticular; pl, palatine; pmx, premaxilla; pof, postorbitofrontal; pr, prootic; prf, prefrontal; ps, parasphenoid; pt, pterygoid; pvc, posterior aperture of the vidian canal q; quadrate; rv, right vomer; sa, surangular; scp, sclerotic plate; smx, septomaxilla; so, supraoccipital; sp, splenial; stp, stapes; VII, IX-XII, exit of cranial nerves.

os écaille dessin holotype +2
Holotype of Alcione elainus.
Fig. 6 of:
Longrich, N. R., Martill, D. M., & Andres, B. (2018). Late Maastrichtian pterosaurs from North Africa and mass extinction of Pterosauria at the Cretaceous-Paleogene boundary. PLoS biology, 16(3), e2001663.
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Original figure legend:
A. elainus FSAC-OB 2, holotype partial skeleton and FSAC-OB 217, metacarpal IV.

(A) Holotype right humerus in anterior view, (B) holotype right ulna and radius in anterior view, respectively, (C) holotype sternum in left lateral view, (D) referred metacarpal IV, (E) holotype, distal end of left metacarpal IV and left scapulocoracoid, and (F) holotype right femur in posterior view. Abbreviations: co, coracoid; cr, cristospine; dc, distal condyle; dpc, deltopectoral crest; ect, ectepicondyle; fh, femoral head; gl, glenoid; gt, greater trochanter; hh, humeral head; hum, humerus; mcIV, metacarpal IV, pc, proximal cotyle; pf, pneumatic foramen; rad, radius; scpr, supracondylar process; ste, sternum; uln, ulna.

Holotype of Alcione elainus. Fig. 6 of: Longrich, N. R., Martill, D. M., & Andres, B. (2018). Late Maastrichtian pterosaurs from North Africa and mass extinction of Pterosauria at the Cretaceous-Paleogene boundary. PLoS biology, 16(3), e2001663. --- Original figure legend: A. elainus FSAC-OB 2, holotype partial skeleton and FSAC-OB 217, metacarpal IV. (A) Holotype right humerus in anterior view, (B) holotype right ulna and radius in anterior view, respectively, (C) holotype sternum in left lateral view, (D) referred metacarpal IV, (E) holotype, distal end of left metacarpal IV and left scapulocoracoid, and (F) holotype right femur in posterior view. Abbreviations: co, coracoid; cr, cristospine; dc, distal condyle; dpc, deltopectoral crest; ect, ectepicondyle; fh, femoral head; gl, glenoid; gt, greater trochanter; hh, humeral head; hum, humerus; mcIV, metacarpal IV, pc, proximal cotyle; pf, pneumatic foramen; rad, radius; scpr, supracondylar process; ste, sternum; uln, ulna.

crête humérus Crétacé Maastrichtien +7
Holotype of Alcione elainus.
Fig. 6 of:
Longrich, N. R., Martill, D. M., & Andres, B. (2018). Late Maastrichtian pterosaurs from North Africa and mass extinction of Pterosauria at the Cretaceous-Paleogene boundary. PLoS biology, 16(3), e2001663.
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Original figure legend:
A. elainus FSAC-OB 2, holotype partial skeleton and FSAC-OB 217, metacarpal IV.

(A) Holotype right humerus in anterior view, (B) holotype right ulna and radius in anterior view, respectively, (C) holotype sternum in left lateral view, (D) referred metacarpal IV, (E) holotype, distal end of left metacarpal IV and left scapulocoracoid, and (F) holotype right femur in posterior view. Abbreviations: co, coracoid; cr, cristospine; dc, distal condyle; dpc, deltopectoral crest; ect, ectepicondyle; fh, femoral head; gl, glenoid; gt, greater trochanter; hh, humeral head; hum, humerus; mcIV, metacarpal IV, pc, proximal cotyle; pf, pneumatic foramen; rad, radius; scpr, supracondylar process; ste, sternum; uln, ulna.

Holotype of Alcione elainus. Fig. 6 of: Longrich, N. R., Martill, D. M., & Andres, B. (2018). Late Maastrichtian pterosaurs from North Africa and mass extinction of Pterosauria at the Cretaceous-Paleogene boundary. PLoS biology, 16(3), e2001663. --- Original figure legend: A. elainus FSAC-OB 2, holotype partial skeleton and FSAC-OB 217, metacarpal IV. (A) Holotype right humerus in anterior view, (B) holotype right ulna and radius in anterior view, respectively, (C) holotype sternum in left lateral view, (D) referred metacarpal IV, (E) holotype, distal end of left metacarpal IV and left scapulocoracoid, and (F) holotype right femur in posterior view. Abbreviations: co, coracoid; cr, cristospine; dc, distal condyle; dpc, deltopectoral crest; ect, ectepicondyle; fh, femoral head; gl, glenoid; gt, greater trochanter; hh, humeral head; hum, humerus; mcIV, metacarpal IV, pc, proximal cotyle; pf, pneumatic foramen; rad, radius; scpr, supracondylar process; ste, sternum; uln, ulna.

crête humérus Crétacé Maastrichtien +7
Various fossils pertaining to the holotype of the Triassic ichthyosaur Toretocnemus (originally Leptocheirus, also Merriamia) zitteli. This image is derived from plate 23 in Merriam (1903), done by an uncredited artist. The arrangement of the individual figures has been modified from the original (most notably, figures 3 and 4 were swapped to minimize confusion).
Original description:
Leptocheirus zitteli n. gen. and sp.
Figures reproduced natural size from the type specimen.

Fig. 1.— Right side of skull.
Fig. 2. — Cross-section of posterior portion of the lower jaw, taken above the point marked A on the lateral view of the skull.
Fig. 3. — Cross-section of the upper and lower jaws, taken at the break immediately behind the point marked Sp on the lateral view of the skull.
Fig. 4. — Lateral view of an anterior caudal centrum.
Legend (modified from original):

Ar - articular
A - angular
D - dentary
J - jugal
L - lacrimal
Mx - maxilla
Po - postorbital
Sa - surangular
Se - sclerotic ring
Sp - spenial
T - cross-section of tooth
X - doubtful element

Various fossils pertaining to the holotype of the Triassic ichthyosaur Toretocnemus (originally Leptocheirus, also Merriamia) zitteli. This image is derived from plate 23 in Merriam (1903), done by an uncredited artist. The arrangement of the individual figures has been modified from the original (most notably, figures 3 and 4 were swapped to minimize confusion). Original description: Leptocheirus zitteli n. gen. and sp. Figures reproduced natural size from the type specimen. Fig. 1.— Right side of skull. Fig. 2. — Cross-section of posterior portion of the lower jaw, taken above the point marked A on the lateral view of the skull. Fig. 3. — Cross-section of the upper and lower jaws, taken at the break immediately behind the point marked Sp on the lateral view of the skull. Fig. 4. — Lateral view of an anterior caudal centrum. Legend (modified from original): Ar - articular A - angular D - dentary J - jugal L - lacrimal Mx - maxilla Po - postorbital Sa - surangular Se - sclerotic ring Sp - spenial T - cross-section of tooth X - doubtful element

description Trias fossile holotype +7
The shoulder girdle of the holotype of the Triassic ichthyosaur Toretocnemus (originally Leptocheirus, also Merriamia) zitteli. This image is derived from plate 21 in Merriam (1903), done by an uncredited artist. The arrangement of the individual figures has been slightly modified from the original.
Original description: Leptocheirus zitteli n. gen. and sp.
Figures reproduced natural size from the type specimen.

Fig. 1.—Clavicles (cl) and probable interelavicle (Ic and Ic')- The interclavicle has been separated into two fragments by a crack in the matrix.
Fig. 2.—Inner side of the coracoids and the left scapula.

The shoulder girdle of the holotype of the Triassic ichthyosaur Toretocnemus (originally Leptocheirus, also Merriamia) zitteli. This image is derived from plate 21 in Merriam (1903), done by an uncredited artist. The arrangement of the individual figures has been slightly modified from the original. Original description: Leptocheirus zitteli n. gen. and sp. Figures reproduced natural size from the type specimen. Fig. 1.—Clavicles (cl) and probable interelavicle (Ic and Ic')- The interclavicle has been separated into two fragments by a crack in the matrix. Fig. 2.—Inner side of the coracoids and the left scapula.

description Trias holotype spécimen +5
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