holotype

Braincase of WSC 10058, holotype of Ornatops Braincase of Ornatops incantatus in digital 3D model in right lateral (A and C) and left lateral (D and F) views, with enlarged images of the right (B) and left (E) auditory regions. Credit: McDonald AT, Wolfe DG, Freedman Fowler EA, Gates TA. 2021. A new brachylophosaurin (Dinosauria: Hadrosauridae) from the Upper Cretaceous Menefee Formation of New Mexico. PeerJ 9:e11084 https://doi.org/10.7717/peerj.11084

The hadrosaurid dinosaur Wulagasaurus dongi gen. et sp. nov. from the Upper Cretaceous Yuliangze Formation at the Wulaga quarry, China. A. Holotype GMH W184, right dentary in lateral (A1) and medial (A2) views. B. GMH W217, right dentary in lateral (B1) and medial (B2) views.

The holotype skeleton of Kamuysaurus japonicus gen. et sp. nov. (b) Reconstructed skeleton, showing recovered elements. Selected postcranial elements: cervical vertebrae (atlas, axis, and fourth and twelfth cervicals) in left lateral view (c), dorsal vertebrae (first, seventh, and sixteenth dorsals) in left lateral view (d), caudal vertebrae (anterior, middle, and posterior caudals) in left lateral view (e), left scapula (f) and coracoid (g) in lateral view, right sternum in ventral view (h), left humerus in anterior view (i), right ulna and radius in medial view (j), right manus in dorsal view (k), right pelvis in lateral view (l), right femur in anterior view (m), right tibia in anterior view (n), right fibula in lateral view (o), right astragalus and calcaneum, articulated positioned with tibia (p), and right pes in dorsal view (q). All scales are 10 cm except 1 m scale for (b). Abbreviations; ac, acromion process; ast, astragalus; cal, calcaneum; dpc, deltopectoral crest; fi, fibula; icg, intercondylar groove; il, ilium; is, ischium; ltr, lesser trochanter; mc2, metacarpal II; mc5, metacarpal V; mt2, metatarsal II; mt4, metatarsal IV; nc, neurocentrum; od, odontoid; olp, olecranon process; pc, pleurocentrum; pu, pubis; ra, radius; ti, tibia; ul, ulna; vp, ventral process.

Graphite drawing with digital retouches of the holotype (UCMP 152028) of Adelolophus hutchisoni. (A) lateral view; (B) medial view; (C) dorsolateral view. The scale bar is equal to 10 centimeters. Reference: Figure 9.2 from Gates et al. (2014).[1]

Holotype maxilla of the lambeosaurine hadrosaurid Ajnabia odysseus from the Late Cretaceous (late Maastrichtian) of Sidi Chennane, Morocco

Right premaxilla of Eotrachodon orientalis (holotype MSC 7949). (A) Dorsal view. (B) Rostral view. (C) Lateral view. (D) Rostral view of rostral layer of premaxillary denticles. (E) Ventral view of premaxillary denticles. (F) Ventral view. (G) Medioventral view.

Photograph of the right dentary of the holotype (MNHAH D1-033516) of the hadrosaurid Yamatosaurus izanagii (Kobayashi et al. 2021). A is the lateral view; B is the medial view; C is the dorsal view; D is the ventral view; E is the anterior view. The white numbers (B) show the positions of nutrient foramina.

Dentary of the hadrosauroid dinosaur Fylax thyrakolasus gen. et sp. nov. (IPS-36338, holotype) from the uppermost Maastrichtian Fontllonga-R locality; in posterior (A1), medial (A2), dorsal (A4), anterior (A5), lateral (A6), and ventral (A7) views. A detailed lingual view of the tooth crowns appears in A3.

Elements of the Qianjiangsaurus changshengi holotype (CLGRP V00016), including the predentary in dorsal view (top left), left dentary in medial view (top right), and a block containing eighteen mostly articulated anterior to middle caudal vertebrae and associated chevrons and ossified tendons.

Left ilium of Cedrorestes crichtoni (DMNH 47994, holotype).

Dorsal vertebrae series of the holotype specimen of Morelladon beltrani (CMP-MS-03). CMP-MS-03-06, -07 (including CMP-MS-03-17 and -29) and -05 (including CMP-MS-03-08) in left lateral (A) view. Interpretive drawing of CMP-MS-03-05 (including CMP-MS-03-08 neural spine) in left lateral (B) view. Abbreviations: ns, neural spine; poz, postzygapophysis; pre, prezygapophysis; rec, vertical recess; tp, transverse process.

Theiophytalia kerri Brill & Carpenter, 2006 - ornithopod dinosaur skull from the Cretaceous of Colorado, USA. (YPM 1887, public display, Garden of the Gods visitor center, Colorado Springs, Colorado, USA) This skull is the holotype and only known specimen of an entire genus of ornithopod dinosaurs, Theiophytalia. Ornithopods were herbivorous dinosaurs. From exhibit signage: A Brand New Dinosaur Species Theiophytalia kerri What we know about the dinosaur fossil - so far: It is the only Theiophytalia kerri fossil known to exist in the world. It is a brand new genus and species of dinosaur. Theiophytalia kerri means "belonging to the Garden of the Gods". "Theios" is a Greek word that means "belonging to the gods" and "phytalia" means "garden". "kerri" honors James Hutchinson Kerr (pronounced "Care"), who discovered the dinosaur fossil. Theiophytalia was a medium-sized dinosaur, measuring about 30 feet from head to tail. It was found in 1878 in the Garden of the Gods in the lower Lytle Member of the Purgatoire Rock Formation of the Dakota Group. The dinosaur fossil skull is from the Aptian-Albian Age of the Cretaceous Ear, 125 - 100 million years old. Look closely at the fossil skull. Where are the teeth located? You can see the teeth in the back of the jaw. There are no teeth in the front of the dinosaur's beak-like mouth. This is evidence that the Theiophytalia was a plant eater. It could nip and tear plants, then grind them between its tightly-packed back teeth, similar to plant-eaters of today. Timeline: Dinosaur Fossil Discovered, then Forgotten 1878 - James Kerr, geology professor at Colorado College, finds a fossil skull "in one of the ridges east of the red rocks of the Garden of the Gods." 1886 - O.C. Marsh, famous 1800s dinosaur collector from Yale University, obtains the fossil skull from James Kerr, identifies it as a Camptosaurus dinosaur, and sends the fossil to the Yale Peabody Museum in New Haven, Connecticut. 1886 - 1995 - While the dinosaur fossil skull safely rests in the Yale Museum for many decades, knowledge of its existence is forgotten in Colorado Springs. Rediscovery - 117 years later 1994 - Colorado Springs City park staff research new exhibits for the Garden of the Gods Visitor and Nature Center, set to open in 1995. They meet with Doctor Kirk Johnson, curator of paleontology at the Denver Museum of Nature & Science. 1995 - Kirk Johnson refers the park staff to his museum colleague Doctor Ken Carpenter, expert dinosaur scientist. Doctor Carpenter remembers seeing in his files "something about a dinosaur fossil found in Garden of the Gods" and mails the following article to park staff. Camptosaurus amplus No. 1887, Yale Museum, consisting of portions of the skull and lower jaw. It was collected from deposits in the Garden of the Gods, Colorado Springs, Colorado. With this specimen was found the following note in Professor O.C. Marsh's handwriting, "Part of this animal and various Sauropoda bones were taken out by Professor Kerr in 1878." A Case of Mistaken Identity Reveals a Brand New Dinosaur! 1996 - Kirk Johnson (a Yale alumnus) secures permission to hand-carry the Camptosaurus fossil from the Yale Peabody Museum to Denver so that Ken Carpenter can make a cast (a precise replica) of the fossil. Doctor Carpenter notices irregularities in the Camptosaurus fossil and decides to re-examine the fossil when his schedule permits. 1997 - The Camptosaurus fossil replica is given to the City of Colorado Springs and is exhibited at the Garden of the Gods Visitor and Nature Center. 2006 - Doctor Carpenter and his associate Kathleen Brill reassess the fossil skull and note that it differs from other Camptosaurus skulls in several significant ways, such as the narrower mouth and snout, and the position of the nasal openings, and the bony structures over the eyes. Also, microscopic identification of the rock matrix clinging to the fossil, and research of archival maps, reveal that the fossil skull was actually found in the lower Dakota Rock Formation, not the Morrison Formation as originally reported in the 1800s. The skull is from a dinosaur that lived in the Cretaceous Ear and can't be a Jurassic Era Camptosaurus. Doctor Carpenter's exacting research reveals that the dinosaur skull is a brand new genus and species of dinosaur! He names it Theiophytalia kerri. 2008 - The Theiophytalia kerri is proudly re-exhibited at the Garden of the Gods Visitor and Nature Center. Classification: Animalia, Chordata, Vertebrata, Dinosauria, Ornithischia, Ornithopoda Stratigraphy: lower Lytle Member, Purgatoire Formation, Dakota Group, Aptian to Albian Stages, upper Lower Cretaceous Locality: Garden of the Gods, Colorado Springs, Colorado, USA See info. at: en.wikipedia.org/wiki/Theiophytalia and en.wikipedia.org/wiki/Ornithopoda

Holotype of Pulaosaurus (IVPP V30936) in left lateral view

Changmiania liaoningensis, an ornithopod dinosaur from the Lower Cretaceous of Lujiatun (Liaoning Province, China). (A) Holotype PMOL AD00114 in dorsal view; (B) anterior part of the holotype PMOL AD00114 in caudolateral view; (C) referred specimen PMOL LFV022 in dorsal view. Red arrows indicate the emplacement of the gastrolith clusters.

Chilesaurus holotype cast.

Holotype of Jakapil kaniukura (MPCA-PV-630), postcranial bones. Speculative silhouette showing preserved elements (a); osteoderm distribution is speculative and partial to show non-osteodermal elements); dorsal vertebra elements in dorsal (b), right lateral (c) and anterior (d,e) views; sacral vertebra in left lateral view (f); mid-caudal vertebra in left lateral view (g); fragment of the mid-shaft of a dorsal rib in posterior view (the enlarged, broken posterior edge is highlighted (h); expanded distal ends of two dorsal ribs (i); left scapula in lateral view (j); right scapula in lateral view (k); right coracoid in lateral view (l); left and right humeri in anterior view (m); probable right ulna in lateral view (n); metacarpals, non-ungual and ungual phalanx in dorsal views (o); left femur elements in anterior view (p); proximal end of the right fibula in lateral view (q); distal end of the left tibia in anterior view (r); ischial elements in side view (s); cervical osteoderms in dorsal view (t), flat scutes in dorsal view (u), spine-like osteoderm in side view (v) and ossicle in dorsal view (w). ac acromial crest, aco asymmetrical cervical osteoderm, alp anterolateral process, ap acromial process, at anterior trochanter, bb basal bone, ebr expanded broken rib edge, di diapophysis, dpc deltopectoral crest, ft fourth trochanter, gl glenoid, mc metacarpals, nc neural canal, ncs neurocentral suture, ph non-ungual phalanx, pp pubic peduncle, poz postzygapophyses, rug marginal rugosities, sb scapular blade, sc scute, tp transverse process, uph ungual phalanx.