formation

Pelvis (fused ilia and pubes) of UALVP 48778, holotype of Hesperonychus elizabethae, from the late Campanian Dinosaur Park Formation of Alberta, Canada. Fossil collected by Dr. Elizabeth Nicholls. Prepared by Clive Coy. Digital photography by Nick Longrich. Additional digital editing by Nick Longrich.

Arcovenator escotae (Theropoda, Abelisauridae), braincase (MHNAix-PV 2011-12) in dorsal view (Lower Argiles Rutilantes Formation, Jas Neuf Sud, Var)

Left ilium of Stokesosaurus clevelandi, Madsen 1974 (UMNH VP 7473), Morrison Formation, Utah, USA, Late Jurassic (early Tithonian).

Tooth of cf. Zapsalis, with close up of denticles. Specimen UALVP 49582 from the Milk River Formation.

Skeletal reconstructions of Dinosaur Park Formation caenagnathids. Skeletal reconstructions of Citipes elegans (left), Chirostenotes pergracilis (middle), and Caenagnathus collinsi (right), showing variation in skeletal representation and body size. Previously referred material is indicated in white and newly referred material is indicated in red for each taxon. Blue asterisks indicate elements that have been histologically sampled for each taxon. Citipes elegans: dentaries, metatarsal IV; Chirostenotes pergracilis: dentaries, tibia; Caenagnathus collinsi: pubis.

Right dentary of Zalmoxes shqiperorum holotype NHMUK R4900in dorsal and lateral views, from the Sinpetru Formation of Sânpetru (Romania; outline based on the specimen UBB NVZ1-1).

The nodosaurid dinosaur, Glyptodontopelta is mostly known from isolated osteoderms ("armor plates"). The specimen SMP VP-1580 (State Museum of Pennysylvania) is however, the most complete known specimen, comprising parts of the skull, hundreds of osteoderms and fragments. This specimen was discovered in 2003 by Warwick Fowler, in the Naashoibito Member of the Ojo Alamo Formation (Maatrichtian, Late Cretaceous) of the San Juan Basin, New Mexico, during an expedition led by Dr. Robert Sullivan.

Drawing of lectoholotype skull BMNH R.3386 (desigated by Weishampel et al., 1993)[1][2] of Limnosaurus transsylvanicus NOPCSA 1900 (later renamed Telmatosaurus transsylvanicus), a basal hadrosaurid dinosaur from the Late Maastrichtian (latest Upper Cretaceous) Sanpetru Formation of West Central Romania.

Right dentary of Zalmoxes robustus holotype NHMUK R3392 in dorsal and lateral views, from the Sinpetru Formation, Sânpetru (Romania)

‘Koutalisaurus kohlerorum’ (Ornithopoda, Lambeosaurinae; indeterminate lambeosaurine sensu Prieto-Márquez et al. 2013), right dentary (IPS 29920 (formerly IPS SRA 27) in medial view (‘lower red unit’ of the Tremp Formation, Les Llaus near Sant Romà d’Abella, Lleida)

Theiophytalia kerri Brill & Carpenter, 2006 - ornithopod dinosaur skull from the Cretaceous of Colorado, USA. (YPM 1887, public display, Garden of the Gods visitor center, Colorado Springs, Colorado, USA) This skull is the holotype and only known specimen of an entire genus of ornithopod dinosaurs, Theiophytalia. Ornithopods were herbivorous dinosaurs. From exhibit signage: A Brand New Dinosaur Species Theiophytalia kerri What we know about the dinosaur fossil - so far: It is the only Theiophytalia kerri fossil known to exist in the world. It is a brand new genus and species of dinosaur. Theiophytalia kerri means "belonging to the Garden of the Gods". "Theios" is a Greek word that means "belonging to the gods" and "phytalia" means "garden". "kerri" honors James Hutchinson Kerr (pronounced "Care"), who discovered the dinosaur fossil. Theiophytalia was a medium-sized dinosaur, measuring about 30 feet from head to tail. It was found in 1878 in the Garden of the Gods in the lower Lytle Member of the Purgatoire Rock Formation of the Dakota Group. The dinosaur fossil skull is from the Aptian-Albian Age of the Cretaceous Ear, 125 - 100 million years old. Look closely at the fossil skull. Where are the teeth located? You can see the teeth in the back of the jaw. There are no teeth in the front of the dinosaur's beak-like mouth. This is evidence that the Theiophytalia was a plant eater. It could nip and tear plants, then grind them between its tightly-packed back teeth, similar to plant-eaters of today. Timeline: Dinosaur Fossil Discovered, then Forgotten 1878 - James Kerr, geology professor at Colorado College, finds a fossil skull "in one of the ridges east of the red rocks of the Garden of the Gods." 1886 - O.C. Marsh, famous 1800s dinosaur collector from Yale University, obtains the fossil skull from James Kerr, identifies it as a Camptosaurus dinosaur, and sends the fossil to the Yale Peabody Museum in New Haven, Connecticut. 1886 - 1995 - While the dinosaur fossil skull safely rests in the Yale Museum for many decades, knowledge of its existence is forgotten in Colorado Springs. Rediscovery - 117 years later 1994 - Colorado Springs City park staff research new exhibits for the Garden of the Gods Visitor and Nature Center, set to open in 1995. They meet with Doctor Kirk Johnson, curator of paleontology at the Denver Museum of Nature & Science. 1995 - Kirk Johnson refers the park staff to his museum colleague Doctor Ken Carpenter, expert dinosaur scientist. Doctor Carpenter remembers seeing in his files "something about a dinosaur fossil found in Garden of the Gods" and mails the following article to park staff. Camptosaurus amplus No. 1887, Yale Museum, consisting of portions of the skull and lower jaw. It was collected from deposits in the Garden of the Gods, Colorado Springs, Colorado. With this specimen was found the following note in Professor O.C. Marsh's handwriting, "Part of this animal and various Sauropoda bones were taken out by Professor Kerr in 1878." A Case of Mistaken Identity Reveals a Brand New Dinosaur! 1996 - Kirk Johnson (a Yale alumnus) secures permission to hand-carry the Camptosaurus fossil from the Yale Peabody Museum to Denver so that Ken Carpenter can make a cast (a precise replica) of the fossil. Doctor Carpenter notices irregularities in the Camptosaurus fossil and decides to re-examine the fossil when his schedule permits. 1997 - The Camptosaurus fossil replica is given to the City of Colorado Springs and is exhibited at the Garden of the Gods Visitor and Nature Center. 2006 - Doctor Carpenter and his associate Kathleen Brill reassess the fossil skull and note that it differs from other Camptosaurus skulls in several significant ways, such as the narrower mouth and snout, and the position of the nasal openings, and the bony structures over the eyes. Also, microscopic identification of the rock matrix clinging to the fossil, and research of archival maps, reveal that the fossil skull was actually found in the lower Dakota Rock Formation, not the Morrison Formation as originally reported in the 1800s. The skull is from a dinosaur that lived in the Cretaceous Ear and can't be a Jurassic Era Camptosaurus. Doctor Carpenter's exacting research reveals that the dinosaur skull is a brand new genus and species of dinosaur! He names it Theiophytalia kerri. 2008 - The Theiophytalia kerri is proudly re-exhibited at the Garden of the Gods Visitor and Nature Center. Classification: Animalia, Chordata, Vertebrata, Dinosauria, Ornithischia, Ornithopoda Stratigraphy: lower Lytle Member, Purgatoire Formation, Dakota Group, Aptian to Albian Stages, upper Lower Cretaceous Locality: Garden of the Gods, Colorado Springs, Colorado, USA See info. at: en.wikipedia.org/wiki/Theiophytalia and en.wikipedia.org/wiki/Ornithopoda

Bakonydraco galaczi gen. et sp. nov.MTMGyn/3, Iharkút, Bakony Mts., Hungary, Csehbánya Formation, Santonian (Upper Cretaceous). Mandible in dorsal (A, B), ventral (C, D), and lateral (E, F) views.

Skeletal diagram featuring the optimal remains of the holotype of Graciliceratops mongoliensis: ZPAL MgD-I/156.[1] Found in the Bayan Shireh Formation, in the original description the remains were referred to the genus Microceratops (now obsolete).[2] However, Sereno in 2000 noted that there was no base for this referral, then, he created a new genus and species for this specimen.[1] The holotype is very fragmented (specially the skull), consisting of:[2] Fragmented skull; 4 cervical, 12 dorsal and 7 sacral vertebrae; right scapula; proximal end of left scapula; left coracoid; right humerus, radius and fragmentary ulna; proximal and distal end of left humerus; proximal fragments of both pubis; fragments of both illium and fragment of right ischium; right femur, tibia and nearly complete pes; distal part of left tibia, fragmentary left pes; tarsals and isolated ribs.[2] The sacral vertebrae are not fused, an indicator of the immaturity of this specimen; the estimated adult size is about 2 meters long or similar to Protoceratops.[2][1] Right quadratojugal, quadrate and fragmentary jugal were reversed in order to get an optimal view.

Aralazhdarcho bostobensis, ZIN PH 57/43, a proximal fragment of a left humerus in proximal (a), ventral (b), anterior (c), dorsal (d), and posterior (e) views. This specimen is from the Shakh Shakh II locality in the northeasten Aral Sea region of Kazakhstan; Bostobe Formation, Upper Cretaceous (Santonian – lower Campanian). Abbreviations: h, humeral head; pf, pneumatic foramen; uc, ulnar crest. Scale bar is 10 mm.

View of the badlands of the type locality for the North Horn Formation on North Horn Mountain.

Cervical vertebrae of rebbachisaurid sauropod Katepensaurus goicoecheai Ibiricu, Casal, Martínez, Lamanna, Luna, and Salgado, 2013a from the Cenomanian–Turonian Bajo Barreal Formation of Chubut Province, Argentina. A. UNPSJB-PV 1007/1, anterior cervical vertebra in anterior (A1) and right ventrolateral (A2) views. B. UNPSJB-PV 1007/2, anterior cervical vertebra in right lateral view. C. UNPSJB-PV 1007/3, middle cervical vertebra in right lateral view. Lateral fossae of the centrum (hypothesized as pneumatic in origin) indicated by arrows.