formation

Dentaries of selected European rhabdodontomorphs in dorsal and lateral views. (A) Obelignathus septimanicus, holotype MDE D30, right dentary from the 'Grès à Reptiles' Formation, Montouliers (France); (B) Mochlodon vorosi holotype MTM V 2010.105.1, left dentary from the Csehbánya Formation, Iharkút (Hungary; mirrored); (C) Mochlodon suessi, lectotype PIUW 2349/2, right dentary from the Grünbach Formation, Muthmannsdorf (Austria); (D) Zalmoxes robustus holotype NHMUK R3392, right dentary from the Sinpetru Formation, Sânpetru (Romania); (E) Rhabdodon priscus, lectotype MPLM 30, left dentary from the Marnes Rouges Inférieures Formation, la Nerthe (France; mirrored drawing after Matheron; outline based on specimens MPLM 30 and MPLM 31, size estimation based on Matheron and Brinkman); (F) Zalmoxes shqiperorum holotype NHMUK R4900, right dentary from the Sinpetru Formation of Sânpetru (Romania; outline based on the specimen UBB NVZ1-1).

Representative taxa from the Santonian Iharkút fauna from the Csehbánya Formation, Bakony Mountains, western Hungary. A Pannoniasaurus inexpectatus (Squamata, Mosasauroidea), dorsal vertebra (MTM uncatalogued) in dorsal view (photo by Réka Kalmár) B Foxemys trabanti (Pleurodira, Bothremydidae), skull (MTM V 2010.215.1.) in dorsal view (photo by Márton Rabi). C Bicuspidon aff. hatzegiensis (Squamata, Borioteiioidea), left dentary (MTM 2006.112.1.) in medial view (photo by László Makádi) D Basal tetanuran (Theropoda, Tetanurae), tooth (MTM V.01.54) in ?lingual view E Indeterminate abelisaurid (Theropoda, Abelisauridae), pedal ungual phalanx (MTM V 2008.43.1.) in lateral view F Pneumatoraptor fodori (Theropoda, Paraves), left scapulocoracoid (holotype, MTM V 2008.38.1.) in lateral view G Mochlodon vorosi (Ornithopoda, Rhabdodontidae), left dentary (holotype, MTM V 2010.105.1) in lateral view H Bakonydraco galaczi (Pterosauria, Azhdarchidae), mandible (holotype, MTM 2007.110.1) in dorsal view I Iharkutosuchus makadii (Eusuchia, Hylaeochampsidae), skull (holotype, MTM 2006.52.1) in dorsal view J Hungarosaurus tormai (Ankylosauria, Nodosauridae), right dentary (MTM 2007.25.2) in lateral view K Bauxitornis mindszentyae (Aves, Enantiornithes), left tarsometatarsus (holotype, MTM V 2009.38.1) in anterior view L Ajkaceratops kozmai (Ceratopsia), fused rostral and premaxillae (holotype, MTM V 2009.192.1) in lateral view. Scale bars: 2 cm in A, V, G, H, I, J; 1 cm in D, E, F, K, L; 1 mm in C.

Representative taxa from the latest Campanian–Maastrichtian faunas from Transylvania, western Romania. A–B Nidophis insularis (Serpentes, Madtsoiidae), articulated vertebrae (LPB (FGGUB) v.547/2) in left lateral (A) and dorsal (B) views (Densuş-Ciula Formation, Tuştea, Haţeg Basin; photo by Ştefan Vasile) C Allodaposuchus precedens (Eusuchia, ?Hylaeochampsidae), skull (PSMUBB V 438) in dorsal view (Sebeş = Şard Formation, Oarda de Jos, southwestern Transylvanian Basin; photo by Vlad Codrea/Massimo Delfino) D Theriosuchus sympiestodon (Mesoeucrocodylia, Atoposauridae), right maxilla (MCDRD 793) in lateral view (Sînpetru Formation, Sînpetru, Haţeg Basin) E–F Indeterminate titanosaur (?Magyarosaurus dacus) (Sauropoda, Titanosauria), isolated osteoderm (LPB (FGGUB) R.1410) in dorsal (E) and lateral (F) views (Sînpetru Formation, Sînpetru, Haţeg Basin) G Indeterminate ornithuran bird (Aves, Ornithurae), incomplete left tibiotarsus (LPB (FGGUB) R.1902) in anterior view (Densuş-Ciula Formation, Vălioara, Haţeg Basin) H Balaur bondoc (Theropoda, Dromaeosauridae), articulated left distal hindlimb (EME PV.313) in lateral view (Sebeş = Şard Formation, Sebeş-Glod, southewestern Transylvanian Basin; photo by Mick Ellison) I Zalmoxes robustus (Ornithopoda, Rhabdodontidae), right dentary (NHMUK R.3407) in medial view (Sînpetru Formation, Sînpetru, Haţeg Basin) J Telmatosaurus transsylvanicus (Hadrosauria), right maxilla (MFGI unnumbered) in lateral view (Sînpetru Formation, Sînpetru, Haţeg Basin) K Indeterminate nodosaurid – Struthiosaurus transylvanicus or new taxon – (Ankylosauria, Nodosauridae), isolated tooth (LPB (FGGUB) R.2182) in medial view (Sînpetru Formation, Sînpetru, Haţeg Basin) L Barbatodon transylvanicus (Multituberculata, Kogaionidae), right maxilla (LPB (FGGUB) M.1635) in medial view (Sînpetru Formation, Pui, Haţeg Basin). Scale bars equal 1 mm in A, B; 5 mm in K; 1 cm in G, L; 2 cm in D; and 5 cm in C, E, F, H, I, J.

Palaeogeographic distribution of late Early and early Late Cretaceous pterosaur assemblages. Taxonomic composition of assemblages shown on Fig. 1. Palaeogeography based on Smith et al. 1994. Abbreviations: 1. Cambridge Greensand, England: 2. Lower Chalk, England: 3. Züümbayan Svita, Khuren-Dukh, Mongolia: 4. Lysaya Gora, Saratov, Russia: 5. Kem Kem red beds, Morocco: 6. Paw Paw Formation, Texas, USA: 7. Lagarcito Formation, San Luis, Argentina: 8. Santana and Crato Formations, Ceara, Brazil: 9. Toolebuc Formation, Queensland, Australia.

Locality map for Australian eurypodan thyreophoran fossils. 1, Stegosaurian? footprint (QM F5701), Walloon Coal Measures, Balgowan Colliery, Balgowan (Bajocian–Bathonian); 2, Minmi paravertebra holotype (QM F10329) (Molnar, 1980), Minmi Member, Bungil Formation (Valanginian–Barremian); 3, Thyreophoran trackways, Broome Sandstone, Dampier Peninsula, Western Australia (Valanginian–Barremian); 4, Ankylosauria indet. (see Barrett et al., 2010) ‘Flat Rocks’ Wonthaggi Formation (upper Hauterivian–Albian); 5, NMV P216739, ‘Lake Copco–Dinosaur Cove’ Eumeralla Formation (middle upper Aptian to lower middle Albian) (Barrett et al., 2010); 6, QM F33286; 7, AM F119849 and AM F35259; 8, Kunbarrasaurus ieversi gen. et sp. nov. (formerly Minmi sp.) (QM F18101); 9, QM F33565 and QM F33566; 10, QM F44324-28. Legend: Dark Green, Toolebuc Formation (late middle–early late Albian); Green, Allaru Formation (upper Albian–(?)lower Cenomanian); Light green, Mackunda Formation (upper Albian–lower Cenomanian); Lightest green, Winton Formation (late Albian–early Turonian).

Original figure caption: .mw-parser-output .smallcaps{font-variant:small-caps}The Middletown Slab covered with the Footprints of Carnivorous Dinosaurs. The tracks are in high relief. Additional notes: Most if not all of these tridactylous (i.e. three-toed) footprints/tracks (but not the actual trackmaker!) are referred to as Grallator or as Grallator-type trace fossils. “High relief” means that these are actually casts of footprints forming a positive relief on the lower surface of the sandstone slab (so-called positive hyporelief). The material that originally formed the mud over which the dinosaurs walked was too friable to be recovered from the quarry in one piece. The slab consists of so called ‘brownstone’ which is the trading name of the sandstone quarried at Middletown, Connecticut. This sandstone belongs to the Lower Jurassic Portland Formation of the Hartford Basin (“Connecticut Valley”) and thus to the upper part of the Newark Supergroup. The trackmakers probably were relatively small ‘primitive’ theropod dinosaurs (coelophysoids) such as Podokesaurus the remains of which were recovered from Lower Jurassic deposits of the Hartford Basin.

A pair of Wendiceratops running through the Oldman formation.

Brachylophosaurus canadensis TMP 1990.104.0001, Oldman Formation, Milk River, Alberta. Original, not cast. At the Royal Tyrrell Museum of Palaeontology.

Brachylophosaurus canadensis TMP 1990.104.0001, Oldman Formation, Milk River, Alberta. Original, not cast. At the Royal Tyrrell Museum of Palaeontology.

Campanian fauna from the Oldman Formation: Wendiceratops, Corythosaurus, Scolosaurus, Chasmosaurus and "Troodon"

Geographic and stratigraphic relationships of the holotype EMK 0012 and the Loki Quarry in northern Montana. (A) Regional relationships between the cross-border paleontological sites in the Oldman and Judith River formations along the Milk River and in Kennedy Coulee in Alberta and Montana. (B) Generalized stratigraphic section in the Kennedy Coulee area modified after Goodwin & Deino (1989) and Rogers, Eberth & Ramezani (2023) with the relationships between the Foremost and Oldman formations in Canada and the Judith River Formation in Montana. Relative placements of important taxa in this area are indicated. Position of 40Ar/39Ar dates originally obtained by Goodwin & Deino (1989) are shown in relation to the new U–Pb CA-ID-TIMS date for KC061517-1 by Ramezani et al. (2022). Bentonite ash beds are only 5 to 7 cm thick so they are exaggerated for clarity. Scale bars delineated in map view are indicated kilometers and in meters stratigraphically.

Brachylophosaurus, Daspletosaurus and Scolosaurus from the Oldman Formation.

Brachylophosaurus canadensis skull (original). From the Oldman Formation, Milk River, Alberta. On display at the Royal Tyrrell Museum, Alberta, Canada.

Brachylophosaurus canadensis (TMP 2005.000.0029), Royal Tyrrell Museum, Drumheller, Alberta, 2025-07-13; cast, Campanian, from the Oldman Formation

Sandstones of the Milk River Formation flank the Milk river at Writing-on-Stone Provincial Park, Alberta.

African locality: 5, Majunga, Maevarano Formation (Campanian): Rahonavis ostromi.