vertebra

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Alvarezsauridae gen. et sp. indet., posterior caudal vertebrae. Dzharakuduk, Uzbekistan; Bissekty Formation, Upper Cretaceous (Turonian). a-e, ZIN PH 2441/16, two vertebrae preserved in articulation, in anterior (a), dorsal (b), posterior (c), lateral (d), and ventral (e) views. f-j, ZIN PH 2442/16, vertebra missing ventral part of the centrum, in dorsal (f), lateral (g), ventral (h), anterior (i), and posterior (j).
Taxa Dzharaonyx

Alvarezsauridae gen. et sp. indet., posterior caudal vertebrae. Dzharakuduk, Uzbekistan; Bissekty Formation, Upper Cretaceous (Turonian). a-e, ZIN PH 2441/16, two vertebrae preserved in articulation, in anterior (a), dorsal (b), posterior (c), lateral (d), and ventral (e) views. f-j, ZIN PH 2442/16, vertebra missing ventral part of the centrum, in dorsal (f), lateral (g), ventral (h), anterior (i), and posterior (j).

vertebra Uzbekistan Bissekty Cretaceous +4
Cervical vertebra of Calamosaurus foxi
Taxa Calamosaurus

Cervical vertebra of Calamosaurus foxi

vertebra Calamosaurus
Tetanuran theropod Cruxicheiros newmanorum gen. et sp. nov. axial vertebrae (WARMS 15771) from the Chipping Norton Limestone Formation,
Bathonian of the United Kingdom. A. Posterior cervical or anterior dorsal vertebra in posterior view. B. Partial middle−posterior dorsal vertebra in right lat−
eral view showing a sagittal cross−section (B1) and in dorsal view (B2). C. Middle−distal caudal vertebra in left lateral (C1) and posterior (C2) views. D. Mid−
dle−posterior dorsal neural arch in anterior (D1), right lateral (D2, D3), and posterior (D4) views. Photographs (A, B1, B2, C1, C2, D1, D2, D4) and line drawing

(D3). Crossed−hatching indicates matrix and grey tone indicates broken bone.
Taxa Cruxicheiros

Tetanuran theropod Cruxicheiros newmanorum gen. et sp. nov. axial vertebrae (WARMS 15771) from the Chipping Norton Limestone Formation, Bathonian of the United Kingdom. A. Posterior cervical or anterior dorsal vertebra in posterior view. B. Partial middle−posterior dorsal vertebra in right lat− eral view showing a sagittal cross−section (B1) and in dorsal view (B2). C. Middle−distal caudal vertebra in left lateral (C1) and posterior (C2) views. D. Mid− dle−posterior dorsal neural arch in anterior (D1), right lateral (D2, D3), and posterior (D4) views. Photographs (A, B1, B2, C1, C2, D1, D2, D4) and line drawing (D3). Crossed−hatching indicates matrix and grey tone indicates broken bone.

bone vertebra drawing United Kingdom +4
Figure 1. Holotype NHMUK PV R 37591 pelvis and vertebrae of P. milnerae gen. et sp. nov. in (a) left lateral view and (b) right lateral view. atr, antitrochanter; bf, brevis fossa; bfr, brevis fossa rim; bs, brevis shelf; dv, dorsal vertebra; iss, ischial shaft; nc, neural canal; no, notch; obf, obturator foramen; poap, postacetabular process; prap, preacetabular process; puf, pubic fenestra; pus, pubic shaft; ras, rib attachment scar; ri, rim; sac, supra-acetabular crest; sv, sacral vertebra.
Taxa Pendraig

Figure 1. Holotype NHMUK PV R 37591 pelvis and vertebrae of P. milnerae gen. et sp. nov. in (a) left lateral view and (b) right lateral view. atr, antitrochanter; bf, brevis fossa; bfr, brevis fossa rim; bs, brevis shelf; dv, dorsal vertebra; iss, ischial shaft; nc, neural canal; no, notch; obf, obturator foramen; poap, postacetabular process; prap, preacetabular process; puf, pubic fenestra; pus, pubic shaft; ras, rib attachment scar; ri, rim; sac, supra-acetabular crest; sv, sacral vertebra.

crest pelvis vertebra holotype +1
Figure 1: Holotype and skeletal reconstruction of Mosaiceratops azumai, gen. et sp. nov (ZMNH M8856). (a) photograph and line drawing of ZMNH M8856; (b) skeletal reconstruction showing preserved elements in white. Scale bar 10 cm. Abbreviations: a, astragalus; boc, basioccipital; c, calcaneum; cav, caudal vertebra; ch, chevron; cv, cervical vertebra; dr, dorsal rib; dv, dorsal vertebra; f, frontal; fem, femur; fl, fibula; h, humerus; il, ilium; is, ischium; L, left; mt, metatarsal; ph, phalanx/phalanges; po, postorbital; R, right; sk, skull; sq, squamosal; t, tibia; td, tendon; ?, undiagnostic remains.
Taxa Mosaiceratops

Figure 1: Holotype and skeletal reconstruction of Mosaiceratops azumai, gen. et sp. nov (ZMNH M8856). (a) photograph and line drawing of ZMNH M8856; (b) skeletal reconstruction showing preserved elements in white. Scale bar 10 cm. Abbreviations: a, astragalus; boc, basioccipital; c, calcaneum; cav, caudal vertebra; ch, chevron; cv, cervical vertebra; dr, dorsal rib; dv, dorsal vertebra; f, frontal; fem, femur; fl, fibula; h, humerus; il, ilium; is, ischium; L, left; mt, metatarsal; ph, phalanx/phalanges; po, postorbital; R, right; sk, skull; sq, squamosal; t, tibia; td, tendon; ?, undiagnostic remains.

humerus scale vertebra drawing +3
Holotype of Jakapil kaniukura (MPCA-PV-630), postcranial bones. Speculative silhouette showing preserved elements (a); osteoderm distribution is speculative and partial to show non-osteodermal elements); dorsal vertebra elements in dorsal (b), right lateral (c) and anterior (d,e) views; sacral vertebra in left lateral view (f); mid-caudal vertebra in left lateral view (g); fragment of the mid-shaft of a dorsal rib in posterior view (the enlarged, broken posterior edge is highlighted (h); expanded distal ends of two dorsal ribs (i); left scapula in lateral view (j); right scapula in lateral view (k); right coracoid in lateral view (l); left and right humeri in anterior view (m); probable right ulna in lateral view (n); metacarpals, non-ungual and ungual phalanx in dorsal views (o); left femur elements in anterior view (p); proximal end of the right fibula in lateral view (q); distal end of the left tibia in anterior view (r); ischial elements in side view (s); cervical osteoderms in dorsal view (t), flat scutes in dorsal view (u), spine-like osteoderm in side view (v) and ossicle in dorsal view (w). ac acromial crest, aco asymmetrical cervical osteoderm, alp anterolateral process, ap acromial process, at anterior trochanter, bb basal bone, ebr expanded broken rib edge, di diapophysis, dpc deltopectoral crest, ft fourth trochanter, gl glenoid, mc metacarpals, nc neural canal, ncs neurocentral suture, ph non-ungual phalanx, pp pubic peduncle, poz postzygapophyses, rug marginal rugosities, sb scapular blade, sc scute, tp transverse process, uph ungual phalanx.
Taxa Jakapil

Holotype of Jakapil kaniukura (MPCA-PV-630), postcranial bones. Speculative silhouette showing preserved elements (a); osteoderm distribution is speculative and partial to show non-osteodermal elements); dorsal vertebra elements in dorsal (b), right lateral (c) and anterior (d,e) views; sacral vertebra in left lateral view (f); mid-caudal vertebra in left lateral view (g); fragment of the mid-shaft of a dorsal rib in posterior view (the enlarged, broken posterior edge is highlighted (h); expanded distal ends of two dorsal ribs (i); left scapula in lateral view (j); right scapula in lateral view (k); right coracoid in lateral view (l); left and right humeri in anterior view (m); probable right ulna in lateral view (n); metacarpals, non-ungual and ungual phalanx in dorsal views (o); left femur elements in anterior view (p); proximal end of the right fibula in lateral view (q); distal end of the left tibia in anterior view (r); ischial elements in side view (s); cervical osteoderms in dorsal view (t), flat scutes in dorsal view (u), spine-like osteoderm in side view (v) and ossicle in dorsal view (w). ac acromial crest, aco asymmetrical cervical osteoderm, alp anterolateral process, ap acromial process, at anterior trochanter, bb basal bone, ebr expanded broken rib edge, di diapophysis, dpc deltopectoral crest, ft fourth trochanter, gl glenoid, mc metacarpals, nc neural canal, ncs neurocentral suture, ph non-ungual phalanx, pp pubic peduncle, poz postzygapophyses, rug marginal rugosities, sb scapular blade, sc scute, tp transverse process, uph ungual phalanx.

bone vertebra holotype Jakapil +1
PLATE X X X I ~ r.
The vertebral column of Anoplosaurus c~rto~ot~s.
Fig. l. Left lateral aspect of third to seventh cervical eentrums, showing how
the bodies of the vertebrae increase in depth from before backward,
and indicating at the upper anterior corner of each an attachment
for a cervical rib.
1 a. Visceral aspect of third (?) cervical vertebra, showing flattened and
rounded under surface, and expansion towards the anterior end.
Fig. 2. Left lateral aspect of fourteenth to eighteenth eentrums of the series,
showing the somewhat compressed appearance of upper parts of the
bodies of the last five dorsal vertebra.
2 a. Visceral aspect of the last dorsal vertebra, 1~o. 18.
2 b. Posterior (?) articular end of the same centrum, No. 18.
3. The sacrum, showing the neural or superior surface, and giving evi-
dence of the great expansion of the spinal cord in the region of the
second and third sacral vertebra. The outlets for the escape of the
sacral nerves at the sides of the centrum are best seen in the first
three vertebrae.
4. Dorsal surface of vertebral end of a sacral rib, showing the massive end
for attachment to the sides of the centrums of the second and third
vertebrm. Its worn upper surface gave attachment to the neural
arch.
5. Left lateral aspect of a sequence of six early caudal vertebrm numbered
25 to 30, showing decreasing depth of the centrum posteriorly, and
diminishing size in the surface for the attachment of the sacral rib
which remains in union with the 30th vertebra. The chevron facet, e,
also diminishes in size.
5 a. Posterior articular surface and facet for the chevron bone of the

vertebra ~No. 26.
Taxa Anoplosaurus

PLATE X X X I ~ r. The vertebral column of Anoplosaurus c~rto~ot~s. Fig. l. Left lateral aspect of third to seventh cervical eentrums, showing how the bodies of the vertebrae increase in depth from before backward, and indicating at the upper anterior corner of each an attachment for a cervical rib. 1 a. Visceral aspect of third (?) cervical vertebra, showing flattened and rounded under surface, and expansion towards the anterior end. Fig. 2. Left lateral aspect of fourteenth to eighteenth eentrums of the series, showing the somewhat compressed appearance of upper parts of the bodies of the last five dorsal vertebra. 2 a. Visceral aspect of the last dorsal vertebra, 1~o. 18. 2 b. Posterior (?) articular end of the same centrum, No. 18. 3. The sacrum, showing the neural or superior surface, and giving evi- dence of the great expansion of the spinal cord in the region of the second and third sacral vertebra. The outlets for the escape of the sacral nerves at the sides of the centrum are best seen in the first three vertebrae. 4. Dorsal surface of vertebral end of a sacral rib, showing the massive end for attachment to the sides of the centrums of the second and third vertebrm. Its worn upper surface gave attachment to the neural arch. 5. Left lateral aspect of a sequence of six early caudal vertebrm numbered 25 to 30, showing decreasing depth of the centrum posteriorly, and diminishing size in the surface for the attachment of the sacral rib which remains in union with the 30th vertebra. The chevron facet, e, also diminishes in size. 5 a. Posterior articular surface and facet for the chevron bone of the vertebra ~No. 26.

vertebra Anoplosaurus
Representative bones of Pulanesaura eocollum. (a) anterior-to-middle cervical vertebrae (BP/1/6199) in left lateral view; (b) holotypic anterior-most dorsal neural arch (BP/1/6882) in left lateral and anterior views; (c) anterior dorsal neural arch (BP/1/6984) in anterior and right lateral views; (d) anterior mid-dorsal neural arch (BP/1/6183) in anterior and right lateral views; (e) middle dorsal neural arch (BP/1/6770) in posterior view; (f) anterior caudal vertebra (BP/1/6646) in right lateral and posterior views; (g) right humerus (BP/1/6193) in anterior view; (h) left ulna (BP/1/6210) in lateral and proximal views; (i) ?left clavicle (BP/1/6752) in dorsal view; (j) left pedal ungual I (BP/1/6186) in proximal and medial views; (k) left tibia (BP/1/6200) in anterior and lateral views; (l) right ischium (reversed) (BP/1/7366) in lateral view. Abbreviations: aidf, anterior infradiapophyseal fossa; ain, anterior incline of the neural spine; ep, epipophysis; hyp, hyposphene; mr, medial ridge; pp, parapophysis; prdl, prezygodiapophyseal lamina; prz, prezygapophyses; rf, radial fossa; spol, spinopostzygapophyseal lamina; vc, ventral convexity. Scale bars equal 5 cm in a-f and i, j; 10 cm in g, h, k, l. Silhouette drawn by BWM Photographs by BWM.
Taxa Pulanesaura

Representative bones of Pulanesaura eocollum. (a) anterior-to-middle cervical vertebrae (BP/1/6199) in left lateral view; (b) holotypic anterior-most dorsal neural arch (BP/1/6882) in left lateral and anterior views; (c) anterior dorsal neural arch (BP/1/6984) in anterior and right lateral views; (d) anterior mid-dorsal neural arch (BP/1/6183) in anterior and right lateral views; (e) middle dorsal neural arch (BP/1/6770) in posterior view; (f) anterior caudal vertebra (BP/1/6646) in right lateral and posterior views; (g) right humerus (BP/1/6193) in anterior view; (h) left ulna (BP/1/6210) in lateral and proximal views; (i) ?left clavicle (BP/1/6752) in dorsal view; (j) left pedal ungual I (BP/1/6186) in proximal and medial views; (k) left tibia (BP/1/6200) in anterior and lateral views; (l) right ischium (reversed) (BP/1/7366) in lateral view. Abbreviations: aidf, anterior infradiapophyseal fossa; ain, anterior incline of the neural spine; ep, epipophysis; hyp, hyposphene; mr, medial ridge; pp, parapophysis; prdl, prezygodiapophyseal lamina; prz, prezygapophyses; rf, radial fossa; spol, spinopostzygapophyseal lamina; vc, ventral convexity. Scale bars equal 5 cm in a-f and i, j; 10 cm in g, h, k, l. Silhouette drawn by BWM Photographs by BWM.

bone humerus vertebra Pulanesaura
Skeletal reconstruction and exemplar skeletal remains of Lingwulong shenqi. Silhouette showing preserved elements (a); middle cervical vertebra in left lateral (b) and anterior (c) views; anterior dorsal vertebra in left lateral (d) and anterior (e) views; posterior dorsal vertebra in lateral view (f); sacrum and ilium in left lateral view (g); anterior caudal vertebra in left lateral (h) and anterior (i) views; right scapulocoracoid in lateral view (j); right humerus in anterior view (k); left pubis in lateral view (l); right ischium in lateral (m) views; right femur in posterior view (n); and right tibia in lateral view (o). Abbreviations: ap, ambiens process; ar, acromial ridge; ip, iliac peduncle; naf, notch anterior to glenoid; np, neural spine; podl, postzygodiapophyseal lamina; ppr, prezygapophyseal process ridge; prp, prezygapophysis; pvf, posteroventral fossa; slf, shallow lateral fossa; spol, spinopostzygapophyseal lamina; sprl, spinoprezygapophyseal lamina; wls, wing-like structure. Scale bars = 100 cm for a and 5 cm for b–o
Taxa Lingwulong

Skeletal reconstruction and exemplar skeletal remains of Lingwulong shenqi. Silhouette showing preserved elements (a); middle cervical vertebra in left lateral (b) and anterior (c) views; anterior dorsal vertebra in left lateral (d) and anterior (e) views; posterior dorsal vertebra in lateral view (f); sacrum and ilium in left lateral view (g); anterior caudal vertebra in left lateral (h) and anterior (i) views; right scapulocoracoid in lateral view (j); right humerus in anterior view (k); left pubis in lateral view (l); right ischium in lateral (m) views; right femur in posterior view (n); and right tibia in lateral view (o). Abbreviations: ap, ambiens process; ar, acromial ridge; ip, iliac peduncle; naf, notch anterior to glenoid; np, neural spine; podl, postzygodiapophyseal lamina; ppr, prezygapophyseal process ridge; prp, prezygapophysis; pvf, posteroventral fossa; slf, shallow lateral fossa; spol, spinopostzygapophyseal lamina; sprl, spinoprezygapophyseal lamina; wls, wing-like structure. Scale bars = 100 cm for a and 5 cm for b–o

humerus scale vertebra Lingwulong
Skeletal elements of Bajadasaurus pronuspinax gen. et sp. nov (MMCh-PV 75). (A–C) Skull roof and braincase in posterior (A), left lateral (B) and right lateral (C) views. (D,E) Left lower jaw in dorsal (D) and medial (E) views. F, Dentaries in anterior view. (G) Pterygoids in ventral view. (H) Left maxilla in medial view. (I) Left lacrimal in lateral view. (J) Left quadratojugal in lateral view. (K,L) Right quadrate in medial (K) and posterior (L) views. (M) Proatlases in dorsal view. (N) Atlantal neurapophyses in anterior view. (O,P), Axis in left lateral (O) and anterior (P) views. (Q,R) Fifth cervical vertebra in left lateral (Q) and anterior (R) views. an, angular; ar, articular; bo, basioccipital; bt, basal tubera; btp, basipterygoid process; ch, ‘chin’ of dentary; cn, cranial nerve; d, dentary; di, diapophysis; f, frontal; fm, foramen magnum; fo, fenestra ovalis; ls, laterosphenoid; met, metotic foramen; mp, medial process; nc, neural canal; ns, neural spine; os, orbitosphenoid; p, parietal; pfo, pneumatic fossa; po, postorbital; pocdf, postzygapophyseal centrodiapophyseal fossa; podl, postzygodiapophyseal lamina; popr, paraoccipital process; poz, postzygapophysis; pra, prearticular; pre, prezygapophysis; prsl, prespinal lamina; ptf, postemporal fenestra; qf, quadrate fossa; rm, replacement maxillary tooth; sa, surangular; sp, splenial; sq, squamosal; stf, supratemporal fenestra; vk, ventral keel; vp, ventral process.
Taxa Bajadasaurus

Skeletal elements of Bajadasaurus pronuspinax gen. et sp. nov (MMCh-PV 75). (A–C) Skull roof and braincase in posterior (A), left lateral (B) and right lateral (C) views. (D,E) Left lower jaw in dorsal (D) and medial (E) views. F, Dentaries in anterior view. (G) Pterygoids in ventral view. (H) Left maxilla in medial view. (I) Left lacrimal in lateral view. (J) Left quadratojugal in lateral view. (K,L) Right quadrate in medial (K) and posterior (L) views. (M) Proatlases in dorsal view. (N) Atlantal neurapophyses in anterior view. (O,P), Axis in left lateral (O) and anterior (P) views. (Q,R) Fifth cervical vertebra in left lateral (Q) and anterior (R) views. an, angular; ar, articular; bo, basioccipital; bt, basal tubera; btp, basipterygoid process; ch, ‘chin’ of dentary; cn, cranial nerve; d, dentary; di, diapophysis; f, frontal; fm, foramen magnum; fo, fenestra ovalis; ls, laterosphenoid; met, metotic foramen; mp, medial process; nc, neural canal; ns, neural spine; os, orbitosphenoid; p, parietal; pfo, pneumatic fossa; po, postorbital; pocdf, postzygapophyseal centrodiapophyseal fossa; podl, postzygodiapophyseal lamina; popr, paraoccipital process; poz, postzygapophysis; pra, prearticular; pre, prezygapophysis; prsl, prespinal lamina; ptf, postemporal fenestra; qf, quadrate fossa; rm, replacement maxillary tooth; sa, surangular; sp, splenial; sq, squamosal; stf, supratemporal fenestra; vk, ventral keel; vp, ventral process.

vertebra Bajadasaurus skull
Cervical vertebrae of rebbachisaurid sauropod Katepensaurus goicoecheai Ibiricu, Casal, Martínez, Lamanna, Luna, and Salgado, 2013a from the
Cenomanian–Turonian Bajo Barreal Formation of Chubut Province, Argentina. A. UNPSJB-PV 1007/1, anterior cervical vertebra in anterior (A1) and
right ventrolateral (A2) views. B. UNPSJB-PV 1007/2, anterior cervical vertebra in right lateral view. C. UNPSJB-PV 1007/3, middle cervical vertebra in

right lateral view. Lateral fossae of the centrum (hypothesized as pneumatic in origin) indicated by arrows.
Taxa Katepensaurus

Cervical vertebrae of rebbachisaurid sauropod Katepensaurus goicoecheai Ibiricu, Casal, Martínez, Lamanna, Luna, and Salgado, 2013a from the Cenomanian–Turonian Bajo Barreal Formation of Chubut Province, Argentina. A. UNPSJB-PV 1007/1, anterior cervical vertebra in anterior (A1) and right ventrolateral (A2) views. B. UNPSJB-PV 1007/2, anterior cervical vertebra in right lateral view. C. UNPSJB-PV 1007/3, middle cervical vertebra in right lateral view. Lateral fossae of the centrum (hypothesized as pneumatic in origin) indicated by arrows.

vertebra Argentina Cenomanian Turonian +3
Fig. 2. Cathartesaura anaerobica gen. et sp. nov.
Posterior cervical vertebra. A, right lateral view;
B, detail of the centrum showing internal cavities.
Abbreviations: acdl, anterior centrodiapophyseal
lamina; al1, accesory lamina 1; al2,
accesory lamina 2; cam, camerae; cpol, centropostzygapophyseal
lamina; di, diapophysis; ns,
neural spine; pa, parapophysis; pcdl, posterior
centrodiapophyseal lamina; podl, postzygodiapophyseal
lamina; poz, postzygapophysis; prdl,
prezy-godiapophyseal lamina; prz, prezygapophysis;
spol, spinopostzygapophyseal lamina;
sprl, spinoprezygapophyseal lamina; spzal,

suprapostzygapophyseal accesory lamina.
Taxa Cathartesaura

Fig. 2. Cathartesaura anaerobica gen. et sp. nov. Posterior cervical vertebra. A, right lateral view; B, detail of the centrum showing internal cavities. Abbreviations: acdl, anterior centrodiapophyseal lamina; al1, accesory lamina 1; al2, accesory lamina 2; cam, camerae; cpol, centropostzygapophyseal lamina; di, diapophysis; ns, neural spine; pa, parapophysis; pcdl, posterior centrodiapophyseal lamina; podl, postzygodiapophyseal lamina; poz, postzygapophysis; prdl, prezy-godiapophyseal lamina; prz, prezygapophysis; spol, spinopostzygapophyseal lamina; sprl, spinoprezygapophyseal lamina; spzal, suprapostzygapophyseal accesory lamina.

vertebra Cathartesaura
Holotype dorsal vertebra of Nopcsasopondylus.
Taxa Nopcsaspondylus

Holotype dorsal vertebra of Nopcsasopondylus.

vertebra holotype Nopcsaspondylus
Vertebra of the rebbachisaurid sauropod Histriasaurus from Croatia
Taxa Histriasaurus

Vertebra of the rebbachisaurid sauropod Histriasaurus from Croatia

vertebra Croatia Histriasaurus Rebbachisauridae
Caudal vertebra of Astrophocaudia SMU 61732 photo from 2014-04-09 PM
Taxa Astrophocaudia

Caudal vertebra of Astrophocaudia SMU 61732 photo from 2014-04-09 PM

vertebra Astrophocaudia
Photograph of the holotype (USNM 538127) vertebra of the sauropod Dzharatitanus (Averianov & Sues, 2021). A is the posterior view; B is the right lateral view; C is the anterior view. Scale bar equals 10 centimetres.
Taxa Dzharatitanis

Photograph of the holotype (USNM 538127) vertebra of the sauropod Dzharatitanus (Averianov & Sues, 2021). A is the posterior view; B is the right lateral view; C is the anterior view. Scale bar equals 10 centimetres.

scale vertebra holotype Dzharatitanis
1 2 3 4 5

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vertebra predator Dinosauria Kank
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vertebra Canada Colombia fossil Dinosauria Ornithomimosauria bird
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vertebra India discovery
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