Dinosauria

Taxon

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Diuqin is an unenlagiine dinosaur from the Late Cretaceous of what is now Argentina. Unenlagiines, known exclusively from South America, are usually classified as dromaeosaurs though this is sometimes debated. Like dromaeosaurs, they were covered in feathers, carnivorous, and had the large sickle-like claw on the inner toe of each foot. Unique to unenlagiines is their elongated snout, suggesting a piscivorous diet. Diuqin was a medium-sized unenlagiine, at about 4 m in length.

Diuqin is an unenlagiine dinosaur from the Late Cretaceous of what is now Argentina. Unenlagiines, known exclusively from South America, are usually classified as dromaeosaurs though this is sometimes debated. Like dromaeosaurs, they were covered in feathers, carnivorous, and had the large sickle-like claw on the inner toe of each foot. Unique to unenlagiines is their elongated snout, suggesting a piscivorous diet. Diuqin was a medium-sized unenlagiine, at about 4 m in length.

claw feather Argentina Cretaceous +5
Figure 1. Evolution of macroecological traits in Dinosauria. Large scale event in dinosaur evolution (a); the origin of dinosaurs (star), hyperthermals (volcano), the earliest fossil Avialae (bird), the earliest fossil angiosperm (flower), the Cretaceous/Palaeogene mass extinction (asteroid). Phylogeny of dinosaurs (b) redrawn from Sereno and adapted to the current consensus and upon which an ancestral state reconstruction of temperature niche (mean annual temperature) after Chiarenza et al. is plotted; Mesozoic palaeogeographies (c) for Triassic (T), Jurassic (J) and Cretaceous (K). Silhouette colours symbolize body mass for each of the taxa represented; information on dietary habits are plotted after Barrett and Zanno & Makovicky; numbers represent clades discussed through this study: 1, Ornithischia; 2, Thyreophora; 3, Ornithopoda; 4, Hadrosauroidea; 5, Marginocephalia; 6, Ceratopsia; 7, Saurischia; 8, Sauropodomorpha; 9, Sauropoda; 10, Theropoda; 11, Ceratosauria; 12, Tetanurae; 13, Coelurosauria; 14, Maniraptoriformes; 15, Maniraptora; 16, Deinonychosauria; 17, Avialae; 18, Ornithothoraces. Palaeogeographies modified from original plots via R package ‘mapast’ using plate models by Scotese.

Figure 1. Evolution of macroecological traits in Dinosauria. Large scale event in dinosaur evolution (a); the origin of dinosaurs (star), hyperthermals (volcano), the earliest fossil Avialae (bird), the earliest fossil angiosperm (flower), the Cretaceous/Palaeogene mass extinction (asteroid). Phylogeny of dinosaurs (b) redrawn from Sereno and adapted to the current consensus and upon which an ancestral state reconstruction of temperature niche (mean annual temperature) after Chiarenza et al. is plotted; Mesozoic palaeogeographies (c) for Triassic (T), Jurassic (J) and Cretaceous (K). Silhouette colours symbolize body mass for each of the taxa represented; information on dietary habits are plotted after Barrett and Zanno & Makovicky; numbers represent clades discussed through this study: 1, Ornithischia; 2, Thyreophora; 3, Ornithopoda; 4, Hadrosauroidea; 5, Marginocephalia; 6, Ceratopsia; 7, Saurischia; 8, Sauropodomorpha; 9, Sauropoda; 10, Theropoda; 11, Ceratosauria; 12, Tetanurae; 13, Coelurosauria; 14, Maniraptoriformes; 15, Maniraptora; 16, Deinonychosauria; 17, Avialae; 18, Ornithothoraces. Palaeogeographies modified from original plots via R package ‘mapast’ using plate models by Scotese.

scale Cretaceous Jurassic Mesozoic +23
Diagram illustrating the "Temporal paradox" in paleontology. First given it's nickname by Alan Feduccia, the paradox is made up by the fact that almost all feathered dinosaurs are dated to have lived millions of years after Archaeopteryx, the oldest bird (late Jurassic, believed to have existed about 150 million years ago). Only a few of the feathered dinosaurs/birdlike dinosaurs are given an older date than Archaeopteryx.

Diagram illustrating the "Temporal paradox" in paleontology. First given it's nickname by Alan Feduccia, the paradox is made up by the fact that almost all feathered dinosaurs are dated to have lived millions of years after Archaeopteryx, the oldest bird (late Jurassic, believed to have existed about 150 million years ago). Only a few of the feathered dinosaurs/birdlike dinosaurs are given an older date than Archaeopteryx.

Jurassic Late Jurassic Archaeopteryx Coelurosauria +2
Montage of eight different representatives of coelurosaurian dinosaurs: (Clockwise from upper left) GIN 100/42 which may represent Citipati or a different oviraptorosaur, Sinosauropteryx prima, Nothronychus mckinleyi, Tyrannosaurus rex, Bambiraptor feinbergi, Passer domesticus, Struthiomimus altus, Microraptor gui. This is a collection of eight different works already found in Wikimedia Commons (see source field below).

Montage of eight different representatives of coelurosaurian dinosaurs: (Clockwise from upper left) GIN 100/42 which may represent Citipati or a different oviraptorosaur, Sinosauropteryx prima, Nothronychus mckinleyi, Tyrannosaurus rex, Bambiraptor feinbergi, Passer domesticus, Struthiomimus altus, Microraptor gui. This is a collection of eight different works already found in Wikimedia Commons (see source field below).

Bambiraptor Citipati Coelurosauria Dinosauria +6
Simplified cladogram of Dinosauria with the distribution of feathers according to the fossil record. Despite its more ancient origin, it
was only in maniraptoriformes that modern-type feathers (pennaceous feathers) have arisen (Based in Xu & Guo, 2009; Clarke,

2013; Godefroit et al., 2013; Han et al., 2014; Koshchowitz et al., 2014).

Simplified cladogram of Dinosauria with the distribution of feathers according to the fossil record. Despite its more ancient origin, it was only in maniraptoriformes that modern-type feathers (pennaceous feathers) have arisen (Based in Xu & Guo, 2009; Clarke, 2013; Godefroit et al., 2013; Han et al., 2014; Koshchowitz et al., 2014).

feather fossil Dinosauria Maniraptoriformes +1
Ареал карнозавров

Ареал карнозавров

Dinosauria Tetanurae
Reconstruction of the theropod dinosaur Tachiraptor admirabilis. Artwork created with graphite and colored pencils. Author: Edwin Chávez "Disfrasaurio".

Reconstruction of the theropod dinosaur Tachiraptor admirabilis. Artwork created with graphite and colored pencils. Author: Edwin Chávez "Disfrasaurio".

Averostra Dinosauria Tachiraptor
Ceratosaurus nasicornis reconstruction. Matches proportions shown in Gregory S. Paul (The Princeton Field Guide to Dinosaurs, 2010, p. 84)

Ceratosaurus nasicornis reconstruction. Matches proportions shown in Gregory S. Paul (The Princeton Field Guide to Dinosaurs, 2010, p. 84)

Averostra Ceratosauria Dinosauria
Sketch-drawing of the vertebrate faunal assemblage of the Lisowice site (modified from Niedźwiedzki)10. (a) Large, theropod-like predatory archosaur (Smok wawelski); (b) large temnospondyl amphibian (Cyclotosaurus sp.); (c) small predatory dinosaurs (Neotheropoda indet.); (d) temnospondyl amphibian (Gerrothorax sp.); (e) small basal crocodylomorph (Crocodylomorpha indet.); (f) small diapsid (Choristodere-like animal); (g) hybodont sharks (Polyacrodus and Hybodus); (h) coelacanth fish; (i) dipnoan fish (Ptychoceratodus sp.); (j) actinopterygian fish; (k) gigantic dicynodont; (l) dinosauriforms or early dinosaurs (Dinosauriformes indet. or Dinosauria indet.); (m) small lepidosauromorphs (Sphenodontia indet.); (n) pterosaurs (Pterosauria indet.); (o) early mammaliaform (Hallautherium sp.).

Sketch-drawing of the vertebrate faunal assemblage of the Lisowice site (modified from Niedźwiedzki)10. (a) Large, theropod-like predatory archosaur (Smok wawelski); (b) large temnospondyl amphibian (Cyclotosaurus sp.); (c) small predatory dinosaurs (Neotheropoda indet.); (d) temnospondyl amphibian (Gerrothorax sp.); (e) small basal crocodylomorph (Crocodylomorpha indet.); (f) small diapsid (Choristodere-like animal); (g) hybodont sharks (Polyacrodus and Hybodus); (h) coelacanth fish; (i) dipnoan fish (Ptychoceratodus sp.); (j) actinopterygian fish; (k) gigantic dicynodont; (l) dinosauriforms or early dinosaurs (Dinosauriformes indet. or Dinosauria indet.); (m) small lepidosauromorphs (Sphenodontia indet.); (n) pterosaurs (Pterosauria indet.); (o) early mammaliaform (Hallautherium sp.).

drawing Dinosauria Neotheropoda Pterosauria
Diagram made to illustrate the debate over the hands of theropod dinosaurs and their alleged descendants, the birds ( Aves ). In the last years, the difference between the hands of theropods and birds have been an important thing when it comes question the theory that birds evolved from dinosaurs, even though some scientists may could refute it. In the diagram, a Neotheropoda ( 1 ), basal tetanurae ( 2 ), a coelurosaurian ( 3 ), the bird (?)Archaeopteryx ( 4 ) and modern bird ( 5 ).
In 1997, birdexpert Alan Feduccia at University of North Carolina discovered that birds develop hands with the digits II, III and IV ( see The Cincinnati Enquirer, 25 - 10 - 1997 ). This is in contrast with the hands of tetanurae, which seems to have the digits I, II and III. This make it almost impossible for dinosaurs and birds to be closely related, according to Feduccia.
Since the discovery by Feduccia, scientific research have came up with a possible explanation to the mystery of the dinosaur - bird hand difference, called The frame shift hypothesis ( see http://scienceblogs.com/tetrapodzoology/2009/06/limusaurus_is_awesome.php ). This hypothesis is based on a discovery which shows that although bird embryos develop the fingers II, III and IV, the genes which is coding for the external appearance of the digits seems to be from the fingers I, II and III. Based on this, scientists belive this: when neotheropods evolved into tetanurae, the losed digit I ( not digit IV, as earlier suggested ). During this process, the genes which coded for how the digits should looks like ( the number of phalanges, for example ) became refurnished ( see the color spots in th upper section in the diagram to understan ). The discovery of Limusaurus has been said to support this theory ( see https://www.livescience.com/animals/090617-dinosaur-hands.html ).
One thing is that may can be used to refute that Limusaurus should support the Frame shift hypothesis is that Limusaurus was a ceratosaurian, and is dated to be much younger than the oldest tetanurae's.
Also, some tetanure´s may had 4 digits' like the Archaeornithomimus ( see number 4 in the diagram ) ( see also http://dml.cmnh.org/1998Oct/msg00443.html and the Allosaurus hand in the image here: http://upload.wikimedia.org/wikipedia/commons/6/69/Allosaurus-mounted.jpg ). If this is the digits I, II, III and IV. If so, it shows that tetanurae had the digits I, II and III, and not II, II and IV, like birds.
For more in this debate, see text section to my picture Raptor-Archaeopteryx-bird hands differens.JPG at http://commons.wikimedia.org/wiki/File:Raptor-Archaeopteryx-bird_hands_differens.JPG.

Diagram made to illustrate the debate over the hands of theropod dinosaurs and their alleged descendants, the birds ( Aves ). In the last years, the difference between the hands of theropods and birds have been an important thing when it comes question the theory that birds evolved from dinosaurs, even though some scientists may could refute it. In the diagram, a Neotheropoda ( 1 ), basal tetanurae ( 2 ), a coelurosaurian ( 3 ), the bird (?)Archaeopteryx ( 4 ) and modern bird ( 5 ). In 1997, birdexpert Alan Feduccia at University of North Carolina discovered that birds develop hands with the digits II, III and IV ( see The Cincinnati Enquirer, 25 - 10 - 1997 ). This is in contrast with the hands of tetanurae, which seems to have the digits I, II and III. This make it almost impossible for dinosaurs and birds to be closely related, according to Feduccia. Since the discovery by Feduccia, scientific research have came up with a possible explanation to the mystery of the dinosaur - bird hand difference, called The frame shift hypothesis ( see http://scienceblogs.com/tetrapodzoology/2009/06/limusaurus_is_awesome.php ). This hypothesis is based on a discovery which shows that although bird embryos develop the fingers II, III and IV, the genes which is coding for the external appearance of the digits seems to be from the fingers I, II and III. Based on this, scientists belive this: when neotheropods evolved into tetanurae, the losed digit I ( not digit IV, as earlier suggested ). During this process, the genes which coded for how the digits should looks like ( the number of phalanges, for example ) became refurnished ( see the color spots in th upper section in the diagram to understan ). The discovery of Limusaurus has been said to support this theory ( see https://www.livescience.com/animals/090617-dinosaur-hands.html ). One thing is that may can be used to refute that Limusaurus should support the Frame shift hypothesis is that Limusaurus was a ceratosaurian, and is dated to be much younger than the oldest tetanurae's. Also, some tetanure´s may had 4 digits' like the Archaeornithomimus ( see number 4 in the diagram ) ( see also http://dml.cmnh.org/1998Oct/msg00443.html and the Allosaurus hand in the image here: http://upload.wikimedia.org/wikipedia/commons/6/69/Allosaurus-mounted.jpg ). If this is the digits I, II, III and IV. If so, it shows that tetanurae had the digits I, II and III, and not II, II and IV, like birds. For more in this debate, see text section to my picture Raptor-Archaeopteryx-bird hands differens.JPG at http://commons.wikimedia.org/wiki/File:Raptor-Archaeopteryx-bird_hands_differens.JPG.

Archaeopteryx Coelurosauria Dinosauria Neotheropoda +3
Cladogram of amniotes. Based on File:Cladogram Amniota A.jpg

Cladogram of amniotes. Based on File:Cladogram Amniota A.jpg

Coelurosauria Dinosauria Maniraptora Ornithischia +2
Cladogram of the former "Reptilia" (red)

Cladogram of the former "Reptilia" (red)

Coelurosauria Dinosauria Maniraptora Ornithischia +2
Dinosaur art by me!

Dinosaur art by me!

Dinosauria
The Chicago specimen of Archaeopteryx (PA 830), a well-preserved fossil highlighting the transitional features between non-avian dinosaurs and birds, housed at the Field Museum of Natural History.

The Chicago specimen of Archaeopteryx (PA 830), a well-preserved fossil highlighting the transitional features between non-avian dinosaurs and birds, housed at the Field Museum of Natural History.

museum fossil specimen Archaeopteryx +2
Life representation of Bicharracosaurus dionidei

Life representation of Bicharracosaurus dionidei

Bicharracosaurus Dinosauria
A badly drew sketch of Brontoholus (a placeholder for when an actual paleoart of Brontotholus is made.

A badly drew sketch of Brontoholus (a placeholder for when an actual paleoart of Brontotholus is made.

paleoart Brontotholus Dinosauria
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News

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scale toy Dinosauria Diplodocia Haolong
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United Kingdom Jurassic tracks Cetiosauria Dinosauria discovery
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bone vertebra description Canada Colombia Cretaceous Late Cretaceous fossil Dinosauria Ornithomimosauria
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Argentina Maastrichtian fossil Dinosauria discovery extinction new species
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Oligocene Dinosauria mammals
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